,2016,36(10): 犃犮狋犪犅狅狋. 犅狅狉犲犪犾. 犗犮犮犻犱犲狀狋. 犛犻狀. : (2016) 犱狅犻 : /j.issn "#$%&' ( )*+,-./01 1, 2,

Size: px

Start display at page:

Download ",2016,36(10): 犃犮狋犪犅狅狋. 犅狅狉犲犪犾. 犗犮犮犻犱犲狀狋. 犛犻狀. : (2016) 犱狅犻 : /j.issn "#$%&' ( )*+,-./01 1, 2,"

勋兢潘
4 years ago
Views:

$, 518035;2 > @ A B C D, ( E E F, G 510275) :HIJ( 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 )KLMNO, PQR2,S34 < J TU 2VWXY 34 Z [ \ ] S ^? _ ` T H I J a b c a b 3 4, ^?_` HIJab 2V, H I J T! + 2 :(1) ^?$

1 ,2016,36(10): 犃犮狋犪犅狅狋. 犅狅狉犲犪犾. 犗犮犮犻犱犲狀狋. 犛犻狀. : (2016) 犱狅犻 : /j.issn "#$%&' ( )*+,-./01 1, 2, 1, 2, 2, 2 (1 )=! >?, ;2 A B C D, ( E E F, G ) :HIJ( 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 )KLMNO, PQR2,S34 < J TU 2VWXY 34 Z [ \ ] S ^? _ ` T H I J a b c a b 3 4, ^?_` HIJab 2V, H I J T! + 2 :(1) ^? _` THIJab O E, 1600m 2 T > Y * 72R, O 45C66, >R E S, T 88.33% (2) H I J a b, 2 2, ; R a T, ( 犛犮犺犲犳犳犾犲狉犪狅犮狋狅狆犺狔犾犪 ) H I J ( 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 ) ( 犘狊狔犮犺狅狋狉犻犪狉狌犫狉犪 ) ( 犐狋犲犪犮犺犻狀犲狀狊犻狊 ) ( 犃狆狅狉狌狊犪犱犻狅犻犮犪 ) a b T R, :53% 38% 28% 23% 2 22% (3), R a 2 ( 犔犻狋狊犲犪狉狅狋狌狀犱犻犳狅犾犻犪 ) O Ra, HIJRa NM T, a b VW HIJRa (4) 34 abrat A &>B&>C &>D &<E &, RaunkiaerT L ; Z c,^?_` THIJab M T R, -2!)T" # A,$ S H I J a b c % & T ' 2, + (,)* +,-, )./ :HIJ;ab 0;R1 ;^?_`;! 203:Q :A 犛狋狉狌犮狋狌狉犲犆犺犪狉犪犮狋犲狉犻狊狋犻犮狊犪狀犱犛狌犮犮犲狊犻狅狀犃狀犪犾狔狊犻狊狅犳犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊犆狅犿犿狌狀犻狋狔犻狀犑犻狀犵狓犻狀犚犲狊犲狉狏狅犻狉犃狉犲犪 LIU Haijun 1,XU Kewang 2,SUN Hongbin 1,HE Qinghua 2,FAN Qiang 2,LIAO Wenbo 2 (1RescueCenterof WildlifeinShenzhen,Shenzhen,Gguangdong518035,China;2 GuangdongProvincialKeyLaboratoryof PlantResources,SchoolofLifeSciences,SunYat senuniversity,guangzhou510275,china) 犃犫狊狋狉犪犮狋 :Theearliestoriginof 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊,animportantrelictspeciesandnationalrareandendan geredplant,couldbetracedtodevonianofpaleozoic.itsevolutionaryprocessissignificanttostudythe palaeofloraandtheoriginandevolutionofferns.inordertoprovidetheoryandpracticebasisforthepro tectionof 犅. 犻狀狊犻犵狀犻狊,weinvestigatedandanalyzedthecharacteristicsoffloristiccomposition,physiogno : ; : :!"#$%( );&'! $%( ); ( )!*+$%( )!:,-.(1978-),/,01, '! *+2! 34 E mail:liunavy@126.com 5678:9:;,01,, <! 34 E mail:lsslwb@mail.sysu.edu.cn

2 10,-., :^?_` HIJab 0<V 2095 my,structure,speciesdiversityandsuccessionof 犅. 犻狀狊犻犵狀犻狊 communityinjingxinreservoirareaof ShenzhenCitybyusingmethodofstandardplot,whichtheresultsshowsasfolowing:(1) 犅. 犻狀狊犻犵狀犻狊 communityinjingxinreservoirareabelongstothesubtropicalevergreenbroad leavedforestswith72spe ciesofvascularplantsbelongingto66generaand45familiesinthe16quadratsof1600 m 2.Exceptfor cosmopolitan,the geographicalelements ofgeneraare predominated bytropicalone,accountingfor 88.33% oftotalofnon worldgenera.(2)theverticalstructureofcommunitycanbedividedintotreelay er(withtwosub layers),shrublayerandherblayer,andthelianaamonglayersareabundant.basedon analysisoftheimportancevalueofdiferentpopulations,itindicatesthatthe 犛犮犺犲犳犳犾犲狉犪狅犮狋狅狆犺狔犾犪, 犅. 犻狀狊犻犵狀犻狊, 犘狊狔犮犺狅狋狉犻犪狉狌犫狉犪, 犐狋犲犪犮犺犻狀犲狀狊犻狊 and 犃狆狅狉狌狊犪犱犻狅犻犮犪 arebelongedtodominantpopulations,and theimportantvaluesare53%,38%,28%,23% and22%,respectively.(3)basedonanalysisofage structure, 犃. 犱犻狅犻犮犪, 犐. 犮犺犻狀犲狀狊犻狊 and 犔犻狋狊犲犪狉狅狋狌狀犱犻犳狅犾犻犪 populationsalbelongtotheincreasingpopula tion,but 犅. 犻狀狊犻犵狀犻狊 populationbelongstothedecliningpopulation.thesepopulationagestructuresindi catedthatthecommunityissituatedwithavigorousdevelopmentstageandthe 犅. 犻狀狊犻犵狀犻狊 populationwil beeliminatedastimegoeson.and(4)thefrequencydistributionlawofthecommunityinthestudyareais A >B >C >D < E,whichisinaccordancewiththeL shapedcurveofraunkiaer.bycomparingwith thoseofotherneighboringregions,the 犅. 犻狀狊犻犵狀犻狊 communityinjingxinreservoirareaofshenzhencity showsthelowerspeciesrichness,whichislikelytoafectedbytheseabreezeandhumanactivities,sothe necessaryinterventionandmeasuresshouldbetakentoprotectthe 犅. 犻狀狊犻犵狀犻狊 communityonsomeextant, forexample,toeliminateupperdeadwood,witheredvine,andstrengthentransparencyunderforest,soas toobtainmoreilumination. 犓犲狔狑狅狉犱狊 : 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 ;community characteristics;species diversity;jingxin reservoir area, ShenzhenCity;protectionstrategy HIJ[ 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 (Hook.)J.Sm.] O J C (Blechnaceae)H I J ( 犅狉犪犻狀犲犪 J. Sm.) [1],, > 2 J > [2] KL 56MN, 7 O> 89, : T a,r,; O W < T " #,A= R >?,@HIJABCD, E T!W [3],S34 <J TU 2VWXY N THIJ >2T - <F,G H I H J K T L E [1] ;OS MT N O[ <' XY T, XY [4] 2P [5], Qa R ST O, N6 U?,VW X2YⅡ&! [6] %,2 Y HIJT34,2+ 34G R, Z > [ R \ ] [7 8] 3 ^ 7 [9] <W [10] _`,abc [11] RT Z HIJa bt 0, S HIJabc ` _` ^?_` HIJab 2V, _` S ' M $T, HIJT! ' 234_] :; "#T^?_` THIJab, O K m, ` T K m +,, V, [12] O 2, 2%; 1 O- 300 m C T O E -, A < T " #, ',1 T 14, T _ T _ 2000mm, 140d, S 80%, O>2T 1 T 2 S, 1 T Y O E E T [13], Y 2@ <=> ^?_`` HI

3 ; J Tab, Z[\]c "#, ` 1600m 2, 16310m 10m T _ "#],U ^ 2cm, 1.5m R T ^(DBH) ; 3 _> 13` 2m 2m T _,"# _> YC 2 TR <, T - M / ab [14], abt, RaT R R 1, < ) V ?@01 (1) : ( 犐犞 )= S 1 犚犃 (%)+ S E 犚犘 (%)+ S 犚犉 (%) 1 >, E ` (2)R1 :abt 1 R R1 2 Pielou c!, " C [15 17] R : 犚 0= 犛 2 Margalef : 犈 =( 犛 -1)/ln 犖 3 R1 : Shannon Wiener (H ): 犎 =- 犘犻 ln 犘 4 Pielou : 犑犛犠 = (- 犘犻 ln 犘犻 )/ln 犛 >TS _> YR T#2,$ ;4 5 > T P i RiT 3 ^ YRT3^ T %; 犖 _> YRT3^ # (AB3CDE< D0 "# ` 1600m 2,& abr 2 ( 1 2), ab B + HIJ ab, >HIJ a R4,G K T 0R, ' HIJ ab "#, ab>y *72R, O45C66 >,J 4C5 5R, \ 2C2 2R,R\39C59 65R, a T 1, > C R_ V\ Y T ab> 5R TC@Y C1C, 8R, #C T2.22%, 3~5RTCY4C, #C T8.89%, A(C4R )*C C2+,-C 3R;.CT#R >#R T29.17% / Y 26 3 C > R, #C T 57.78% C, Y1, E - TV 0 T; >, C A(C C +,- C E, E,, XY TE abr T + 2 / [18 19] T>2R\ T c, Table1 1 "#$%&' (B 0F 3G Areal typesofgeneraofseedplantsin 犅. 犻狀狊犻犵狀犻狊 communityinjingxinreservoirareaofshenzhencity Areal type Numberofgenera Ratiooccupiednon worldgenera/% 1. Cosmopolitan E Pantropic E L2E 8L TropicalAsian &tropicalamericandisjuncted E Oldworldtropic E LHE A L TropicalAsia&tropicalOceania E LHE L TropicalAsiatotropicalAfrica E L TropicalAsia Northtemperate L EastAsian & northamericandisjuncted EastAsia >2 Y EndemictoChina : Total

4 10,-., :^?_` HIJab 0<V 2097 Table2 2 "#$%&' ( HIB A?J+, Quantitativecharacteristicsofdominantpopulationsin 犅. 犻狀狊犻犵狀犻狊 communityinjingxinreservoir areaofshenzhencity R Species #` Totalareaat breastheight S1 Relative abundance S Relative frequency S E Relative dominance Important value 犛犮犺犲犳犳犾犲狉犪狅犮狋狅狆犺狔犾犪 HIJ 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊犘狊狔犮犺狅狋狉犻犪狉狌犫狉犪犐狋犲犪犮犺犻狀犲狀狊犻狊犃狆狅狉狌狊犪犱犻狅犻犮犪犔犻狋狊犲犪狉狅狋狌狀犱犻犳狅犾犻犪 A 犌狅狉犱狅狀犻犪犪狓犻犾犪狉犻狊犛犪狆犻狌犿犱犻狊犮狅犾狅狉犇犻狅狊狆狔狉狅狊犿狅狉狉犻狊犻犪狀犪 < 犛狔狕狔犵犻狌犿犺犪狀犮犲犻 =* 犃犮狉狅狀狔犮犺犻犪狆犲犱狌狀犮狌犾犪狋犪 >? 犃犱犻狀犪狀犱狉犪犿犻犾犲狋犻犃狇狌犻犾犪狉犻犪狊犻狀犲狀狊犻狊 ABC\ 犌犾狅犮犺犻犱犻狅狀狑狉犻犵犺狋犻 DE 犈犾犪犲狅犮犪狉狆狌狊犱犲犮犻狆犻犲狀狊犕犪犮犺犻犾狌狊狏犲犾狌狋犻狀犪犐犾犲狓狆狌犫犲狊犮犲狀狊 > DE 犈犾犪犲狅犮犪狉狆狌狊犮犺犻狀犲狀狊犻狊犆狉犪狋狅狓狔犾狌犿犮狅犮犺犻狀犮犺犻狀犲狀狊犲 FG 犕犪犮犺犻犾狌狊狆犪狌犺狅犻 HI 犞犻犫狌狉狀狌犿狊犲犿狆犲狉狏犻狉犲狀狊 J 犆狌狀狀犻狀犵犺犪犿犻犪犾犪狀犮犲狅犾犪狋犪犛狔犿狆犾狅犮狅狊犾犪狀犮犻犳狅犾犻犪 LM 犇犻狆犾狅狊狆狅狉犪犱狌犫犻犪 N 犎狅犿犪犾犻狌犿犮狅犮犺犻狀犮犺犻狀犲狀狊犲 O 犜狅狓犻犮狅犱犲狀犱狉狅狀狊狌犮犮犲犱犪狀犲狌犿 : [>( X 0.01 CRT 0 Note:Thequantitativecharacteristicsofpopulationswhichimportantvalueisbelow0.01areomited. 1 ^?_` HIJabTR ( 1)R\61 113, 7 E K 1, 16, N ( 犎狅犿犪犾犻狌犿 ) ( 犛狋犲狉犮狌犾犻犪 ( 犛狔犿狆犾狅犮狅狊 ) ( 犐犾犲狓 )2 犇犪犾犫犲狉犵犻犪, # T26.23%; E L H E A L, 15, PQ ( 犇犲狊犿狅狊 R S ( 犎犲犾犻犮犻犪 ) 3T U ( 犈狏狅犱犻犪 ) ( 犃狆狅狉狌狊犪 )2 D E ( 犈犾犪犲狅犮犪狉狆狌狊 );E L TG 10, ( 犕犪犮犺犻犾狌狊 ) V ( 犇犪狆犺狀犻狆犺狔犾狌犿 ) - ( 犆狉犪狋狅狓狔犾狌犿 ( 犃狇狌犻犾犪狉犻犪 )2@ Fig.1 Thecompositionofspeciesin 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 communityinjingxinreservoirareaofshenzhencity ( 犌狅狉犱狅狀犻犪 ) E L2E 8L Y W \ 犔犻狋狊犲犪 ;> 2 Y J ( 犆狌狀

5 ; ninghamia) X^,61 >, O T 1 (5 6 ),E T 53, T 88.33%, O Y 7, 11.67% M, ab>e T S, GY R T + E T a b :, O E [13,20 21] 2.2 (AKL)* HIJ T^?_` ab,g E, T R RY < ( 犛狔狕狔犵犻狌犿犺犪狀犮犲犻 ( 犛犪狆犻狌犿犱犻狊犮狅犾狅狉 ), Tb R Y 9 ( 犇犻狅狊狆狔狉狅狊犿狅狉狉犻狊犻犪狀犪 ) a b,u,zy T W abta 8~13 m,k 15.0 m & abt = T, [ ab <;( 2 T, 4 \,2]^ N_ ` 3, 10~15 m, R1a: by = *( 犃犮狉狅狀狔犮犺犻犪狆犲犱狌狀犮狌犾犪狋犪 ) F G ( 犕犪犮犺犻犾狌狊狆犪狌犺狅犻 ) D E ( 犈犾犪犲狅犮犪狉狆狌狊犱犲犮犻狆犻犲狀狊 * ( 犃狇狌犻犾犪狉犻犪狊犻狀犲狀狊犻狊 K ( 犛狔犿狆犾狅犮狅狊犾犪狀犮犻犳狅犾犻犪 )2 J ( 犆狌狀狀犻狀犵犺犪犿犻犪犾犪狀犮犲狅犾犪狋犪 ) 143 R c2 6~10m, - ( 犆狉犪狋狅狓狔犾狌犿犮狅犮犺犻狀犮犺犻狀犲狀狊犲 ) A ( 犌狅狉犱狅狀犻犪犪狓犻犾犪狉犻狊 ( 犛犪狆犻狌犿犱犻狊犮狅犾狅狉 ) <, Y33 R, R 1, T> ab E,^N X VTE ; TX^, ;OA O-,- <T"# A, Y O20m TA T 1~5m, ; RC& 2 R,R 45 R, YHI J ( 犐犾犲狓狆狌犫犲狊犮犲狀狊 ) A B 犳犾犪犫犲犾狌犾犪狋狌犿 ) J( 犅犾犲犮犺狀狌犿狅狉犻犲狀狋犪犾犲 ) ( 犌犪犺狀犻犪狋狉犻狊狋犻狊 ) G abt =, Y 6 R,1 a 9 (( 犌狀犲狋狌犿犾狅犳狌犲狀狊犲 ) (( 犃犾狔狓犻犪狊犻狀犲狀狊犻狊 ) (( 犜犲狋狉犪犮犲狉犪犪狊犻犪狋犻犮犪 ) ( ( 犇犲狀犱狉狅狋狉狅狆犺犲犳狉狌狋犲狊犮犲狀狊 ) H\( 犆狅狆狋狅狊犪狆犲犾狋犪犱犻犳犳狌狊犪 ) 2.3 HIB AM N01 & _"#, X > RaT #` S1 S S E 2 ( 2),, ab>t K, <, T R HI J T 18%, T a R _ > 6 H I J 143 A= RT 5%, X abt 1 ',6HIJ 2Y &!, 2Y!, R, abt 0R 2.4 O 01 XRaT3^ ab> T, A,, a bt < ' [22 24] : Raunkiaer [25] T & ],[ H I J ab R T A & (1% ~20%) B & (21%~40%) C & (41% ~60%) D & (61% ~ 80%)2 E &(81%~100%) 3 &, 2 >, O A & T R K 1, Y 39 R, #R T49.23%, ab>a= R,, ; O B & T Y 11 R, #R T15.38%; O C &TY13R, # R T18.48%; O D & T K R, Y 3 R,@ #R T4.61%; O E & T Y 9 R, # R T 12.31% ; 2 M, A> C\ ( 犌犾狅犮犺犻犱犻狅狀狑狉犻犵犺狋犻 ) L M ( 犇犻狆犾狅狊狆狅狉犪犱狌犫犻犪 ), R 2 Y T TR ab 3!, T ab 2 T " # A,R S R, ; \ 2J, J( 犔犻狀犱狊犪犲犪狅狉犫犻犮狌犾犪狋犪 ) ( 犇犻犮狉犪狀狅狆狋犲狉犻狊犱犻犮犺狅狋狅犿犪 ) I J ( 犃犱犻犪狀狋狌犿 2 HIJab Fig.2 Frequencydiagramof 犅. 犻狀狊犻犵狀犻狊 community

6 10,-., :^?_` HIJab 0<V 2099 B>C>D<E, : RaunkiaerT L, ^?_`` HIJab 3 Tab, ab+tr & Raunkiaer [25] T,E & R ab>t ar2 R,E &,ab5, G, B& C & D & R T,5 ab R, T HIJab>,E & RT O Raunkiaer [25] T (16%),B&2 C &T O T (B &:14%;C &:9%),D & T O,# ^ D B & C & 2 D & T O, ab, # ^ S T 2.5 )* RaV 2,b ab>t7 2 [24] HIJab>, & RaT,RaT Ⅳ& [24],X^ & :I&,DBH < 2.5cm;Ⅱ &,2.5cm DBH< 7.5cm;Ⅲ&,7.5cm DBH<22.5cm;Ⅳ&,DBH 22.5cm & ab>trat R2 0R( HIJ )c XRaT ; 3 M,HIJRa S,Ⅲ&A Y T, C T G, ZY ST2 T C,HIJ, YA T W, Y, 3 T Ra 2 Ⅱ&, Y TⅠ&,Ⅲ &A,ZY, O, Y c )Ra, Ra Ⅲ&A T K,Ⅱ& 4, O ` 3Ra NM S, Ra, 1, Ra,,Sab AT"#, RMN T, c Ⅱ& 2Ⅲ&A S, O Ra <Ⅰ& 2Ⅳ& Y AT, Ⅱ& ZY, D, <Y 3 T,, NM T 9 RaTⅢ& A 2Ⅳ& Y T, NM T, Ra ab> 20% T Ra>6XHIJ, ZY NM T, O V > Tab ;O O^?_``,1R ' T"#, 0R HIJT, 2 RaT c T A, T2Y!H IJ, V 2.6 B 01 PQ abtr < T, Xab T, R1,R1 2 Xab T [13] R 1, Ta b, 1 2 [11] : X^?_` HIJabTR Mar galef Shannon Wiener H 2 Pielou, ; X ab 3 "#%&' ( RHIB A )* Table3 Populationagestructureofdiferentdominantspeciesof 犅. 犻狀狊犻犵狀犻狊 communityinjingxinreservoir areaofshenzhencity R Species ^&3^ PercentageofindividualindiferentDBHclasses/% Ⅰ Ⅱ Ⅲ Ⅳ HIJ 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊犛犮犺犲犳犳犾犲狉犪狅犮狋狅狆犺狔犾犪犃狆狅狉狌狊犪犱犻狅犻犮犪犐狋犲犪犮犺犻狀犲狀狊犻狊犔犻狋狊犲犪狉狅狋狌狀犱犻犳狅犾犻犪犘狊狔犮犺狅狋狉犻犪狉狌犫狉犪 < 犛狔狕狔犵犻狌犿犺犪狀犮犲犻犇犻狅狊狆狔狉狅狊犿狅狉狉犻狊犻犪狀犪

7 ; 4 %&' ((STU) VW ( B X?APQ Table4 Comparisonofdiversityindexof 犅. 犻狀狊犻犵狀犻狊 communityinjingxinreservoirareawiththeothercommunities intheneighboringregions ab Communitytype ` Size/m 2 犚 0 犎 : ab 犆狉狔狆狋狅犮犪狉狔犪犮犺犻狀犲狀狊犻狊 communityindinghushan [20] *?ab 犈狀犱狅狊狆犲狉犿狌犿犮犺犻狀犲狀狊犲 communityinhongkong ab 犛狋犲狉犮狌犾犻犪犾犪狀犮犲狅犾犪狋犪 communityinnanshan,shenzhen [21] A * <ab 犛狔狕狔犵犻狌犿狅犱狅狉犪狋狌犿 communityindapengpenin sula,shenzhen ^?_` HIJab 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 communityinjingxinreservoir area,shenzhen [13] : Thisarticle 5 %&' "#VW< ( B X?APQ Table5 Comparisonofdiversityindexof 犅. 犻狀狊犻犵狀犻狊 communityinjingxinreservoirareawiththeneighboring regionsinshenzhencity Sampleplot ` Size/m 2 犚 0 犎 : " MaluanshanCountryPark PaiyashanNatureReserve TiantoushanNatureReserve [11] '! GuzhaiNatureReserve,Huicheng Jingxinreservoirarea,Tiantoushan : Thisarticle :. HIJab T =,^?_`, BYHIJ ab;. '! HIJabT Shannon Wiener m 2 _T ;. >,Y` THIJab O! =,^?_` THIJab O = Note:.Thelocationof 犅. 犻狀狊犻犵狀犻狊 communityintheeastoftiantoushannaturereserveisdiferentfromjingxinreservoirarea,which bothalconservatedtheevergreenbroadleavedforestcommunities;.theshannon WienerindexofGuzhainaturereserve,Huichengdis trictisaveragevalueof51200m 2 Sampleareas;.Therearetwo 犅. 犻狀狊犻犵狀犻狊 communitiesinthistable.the 犅. 犻狀狊犻犵狀犻狊 communityin Tiantoushannaturereserveislocatedinitsnortheastarea,whilethe 犅. 犻狀狊犻犵狀犻狊 communityinjingxinreservoirareaislocatedinitssouthwest. T Shannon Wiener H, T a b X ( 4): H IJab T1 < 43, S 4 ab, ', ;O 2 3ab O-,-T" # 7,,a b R a A, 1 C, ^?_``!) 7,V1 ' A, ', S ^?_`` TVc %&T!2 & THIJab TR1 T ( 5),^? _ ` ` H I J a b T R < Shannon Wiener K >,6^?_``, THIJa b O " 8'! +,R1 X T!, abt 2R ab by T, R1 _, G ^?_`` HIJab$ T < T! 3 S 34, ^N T T ^4 [27] H I J 2 Y &!,XY TC3 2P S HIJab"# N Y *45C66 72R, T + E S, ab O E ;^?_` HIJabT < 0T34 M, aba^ OV >, V HIJab >T 0 M> ab> S, T R 4, R 0 OE E [28] H I J a b >, >C T 2, C

8 10,-., :^?_` HIJab 0<V 2101 A,ab[> c V HIJabTR1 S 3Z, >, 1 T ab YO[T*+2! ',* < :+Y T! HIJabT 2 ^?_` HIJRaT M,HIJRa 3 T R a,)4 X3 "# > N 1HIJ N, % HIJb ab T ar, Ra, V > R<M <T"# A 22 3 M ab> R 2 Y T, a b > 20% TRa6XHIJ, ZY NM T, T RaT, abt V, SHIJRaT G ) H I J O A 1 MTR, ' T,. [9], ATabM OHIJT ', T +%& += < % & + ", c +, %&#6, _` HIJ ) Y T. < S $T, _` RT S%&, ' SHIJRaT"# 6'4,$ )AHIJabT"# < 7 T34 S ab ( ),5 *+ RaT ),1aRa ab ' *!,, < N H I J - T N _,<. HIJab, $ ', - ;O / 01T"#, 34 ' Sab1 U 7,$' A,abR1 [29] S D R ( T' $, R C$) ',, 23( 犕犻犽犪狀犻犪犿犻犮狉犪狀狋犺犪 )6 M N O a b T 3, ( -, 4 ( 犔狔犵狅犱犻狌犿犿犻犮狉狅狆犺狔犾狌犿 ) 5 R * 6 ( 犜狅犱犱犪犾犻犪犪狊犻犪狋犻犮犪 ) (( 犅狔狋狀犲狉犻犪犵狉犪狀犱犻犳狅犾犻犪 ) Y A T,$ '! +c ', R $)* 2*+ Y :,!"#$$%&' (,)*+,-./ ,89:;! 4: [1] -.>27,c ; c [M]. 8:C M 9 :,1999: [2] 9:;,;. J T [J].E E,1994,2(3):1 11. LIAO W B,ZHANG H D.ThecharacteristicsofPterido phyteflorafrom GuangdongProvince[J]. 犑狅狌狉狀犪犾狅犳犜狉狅狆犻犮犪犾犪狀犱犛狌犫狋狉狅狆犻犮犪犾犅狅狋犪狀狔,1994,2(3):1 11. [3] c,; L,,.2Y! T [J]. < A,2003,24 (1): WANG YJ,ZHANGSZ,LIY, 犲狋, 犪犾.Thefloristiccharac teristicsanddistributionofnationalkeyprotectedwildplants inshenzhen [J]. 犑狅狌狉狀犪犾狅犳犛狅狌狋犺犆犺犻狀犪犃犵狉犻犮狌犾狋狌狉犪犾犝狀犻狏犲狉狊犻狋狔 (NaturalScienceEdition),2003,24(1): [4] = J [J].,2008,(5): OUYANGCJ,WANGZ,LUOYY, 犲狋, 犪犾.Somewildorna mentalfernsinsouthchina[j]. 犌狌犪狀犵犱狅狀犵犔犪狀犱狊犮犪狆犲犃狉犮犺犻狋犲犮狋狌狉犲,2008,(5): [5] B C,D:E, >,.HIJ1 T < [J]. GA(' C9),2008,7(6): FAN Y M,TAO W Q,GAOSZ, 犲狋, 犪犾.Purificationanda nalysisofthe 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 (Hook.)J.Sm.polysaccha rides[j]. 犑狅狌狉狀犪犾狅犳犌狌犪狀犵狕犺狅狌犝狀犻狏犲狉狊犻狋狔 (NaturalScience Edition),2008,7(6): [6] R,. [M].8:CM9:, 2003:17. [7] FG.HIJ[R \H [J].,1992,(2):37. HUANGBQ.Introductionandpotingexperimentof 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 [J]. 犌狌犪狀犵犱狅狀犵犔犪狀犱狊犮犪狆犲犃狉犮犺犻狋犲犮狋狌狉犲,1992, (2):37. [8] IJK.HIJT <7 \][J].>2GL M,1998, (8):19.

9 ; ZENGSJ.Ornamentandbreedingcultivationof 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 [ 犑 ]. 犆犺犻狀犪犉犾狅狑犲狉牔犘犲狀犼犻狀犵,1998,(8):19. [9] N O,3,,.HIJ \^ T34[J]. E E,2008,16(2): GUOJR,WU H,CHEN X, 犲狋, 犪犾.Studiesonthedevelop mentofgametophytein 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 [J]. 犑狅狌狉狀犪犾狅犳犜狉狅狆犻犮犪犾犪狀犱犛狌犫狋狉狅狆犻犮犪犾犅狅狋犪狀狔,2008,16(2): [10] _F@,>,>,.HIJTW [J].F 34,2008,30(6): FANG Y S,YANG Y B,YANG M H, 犲狋, 犪犾.Chemical constituentsfrom thefern 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 (Blechnaceae) [J]. 犃犮狋犪犅狅狋犪狀犻犮犪犢狌狀狀犪狀犻犮犪,2008,30(6): [11] < H IJab 0 [J]. M,2010,19(4): XU X H,WANG X Q,SUN YJ, 犲狋, 犪犾.Analysisoncom munitycharacteristicsof 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 in Maluanshanand itsadjacent[j]. 犑狅狌狉狀犪犾狅犳犘犾犪狀狋犚犲狊狅狌狉犮犲狊犪狀犱犈狀狏犻狉狅狀犿犲狀狋,2010,19(4): [12] P.A Q _ _R34[J]._ _,2009,20(5): SHEN L.Research onthe Branch Lineof WaterSupply SchemeinDapengPeninsulaofShenzhenCity[J]. 犑狅狌狉狀犪犾狅犳犠犪狋犲狉犚犲狊狅狌狉犮犲狊牔犠犪狋犲狉犈狀犵犻狀犲犲狉犻狀犵,2009,20 (5): [13] ; S, T,U,. A _ * <ab 0<R1 34[J].,2006,27(4): ZHANG YX,CHEN HF,QINXS, 犲狋, 犪犾.Characteristics andspeciesdversity of 犛狔狕狔犵犻狌犿狅犱狅狉犪狋狌犿 communityin DapengPninsula,Shenzhen[J]. 犌狌犻犺犪犻犪,2006,27(4): [14] VW, C., R.ab [M]. G: M9:,1996: [15] SIMPSON E H.Measurementofdiversity[J]. 犖犪狋狌狉犲, 1949,163:688. [16] X. ab1 T _ ][M]. 8:> 2 C Y M9:,1994: [17] YOU MS.Developmentofstudiesonthecommunitydiversi ty [J]. 犑狅狌狉狀犪犾狅犳犉狌犼犻犪狀犃犵狉犻犮狌犾狋狌狉犪犾犝狀犻狏犲狉狊犻狋狔,1997, 27(4): [18] 304.>2R\ T [J].F 3 4, 1991,( IV): WUZY.Theareal typesofchinageneraofseedplants[j]. 犃犮狋犪犅狅狋犪狀犻犮犪犢狌狀狀犪狀犻犮犪,1991,(SupplementIV): [19] 304. >2R\ T T 2 [J]. F 34,1993,( IV): WUZY.Theareal typesofchinageneraofseedplants[j]. 犃犮狋犪犅狅狋犪狀犻犮犪犢狌狀狀犪狀犻犮犪,1993,(SupplementIV): [20] R, V a b I. R 1 [J]. C,1983,(1): PENGSL,WANGBX.Analysisontheforestcommunities ofdinghushani.speciesdiversity[j]. 犈犮狅犾狅犵犻犮犪犾犛犮犻犲狀犮犲, 1983,(1): [21] Z[,\]^, a b 1 <V N [J].,2003,23(7): WANG DB,JISY,CHENPF, 犲狋, 犪犾.Astudyonthespe ciesdiversityandsuccessionsituationofnaturalforestcom munitiesinnanshandistrict,shenzhencity[j]. 犃犮狋犪犈犮狅犾狅犵犻犮犪犛犻狀犻犮犪,2003,23(7): [22] N _,P `,3, a,.` J a b T < [J]. M,2007,16(3): GUO W,SHEN RJ,WUJH, 犲狋, 犪犾.Analysisoncommu nitycompositionandstructureof 犘狊犲狌犱狅狋狊狌犵犪犵犪狌狊狊犲狀犻 in Sanqing MountainofJiangxiProvince[J]. 犑狅狌狉狀犪犾狅犳犘犾犪狀狋犚犲狊狅狌狉犮犲狊犪狀犱犈狀狏犻狉狅狀犿犲狀狋,2007,16(3): [23] 0. [M].8: M9:,2000:127. [24] c,i :,. 6? a b 0 < R 1 34[J]. <C,2010,38(6): ZHAOSS,ZENG Q W,XING F W.Characteristicsand speciesdiversityof 犕犪犮犪狉犪狀犵犪狋犪狀犪狉犻狌狊 communityonilha Verde Hilof Macau[J]. 犑狅狌狉狀犪犾狅犳犃狀犺狌犻犃犵狉犻. 犛犮犻., 2010,38(6): [25] RAUNKIAER C.TheLifeFormsofPlantsandStatistical PlantGeography[M].Oxford:ClarendonPress,1934:632. [26],3, _,. '! H I Jab 034[J]. <CY,2014,30(5):7 12. CAIZ W,WU LF,CHENGF, 犲狋犪犾.Studyof 犅狉犪犻狀犲犪犻狀狊犻犵狀犻狊 communities characteristics at Guzhai Natural Re serve,huichengdistrict[j]. 犉狅狉犲狊狋狉狔犛犮犻犲狀犮犲犪狀犱犜犲犮犺狀狅犾狅犵狔狅犳犌狌犪狀犵犱狅狀犵犘狉狅狏犻狀犮犲,2014,30(5):7 12. [27] <, `, < S [J].,2002,22(3): CHULY,NINGSJ,TANGRQ.Therareandendangered plantsandtheirconservationpoliciesinjiuwan Mountainof Guangxi[J]. 犌狌犻犺犪犻犪,2002,22(3): [28],.- E [M]. G: M9:,1992: [29], -, R. T a b [J].,2001,21(3): ZHOU H C,REN H,PENGSL.Communitystructureof threesecondaryforestsin Nan aoisland,guangdong[j]. 犌狌犻犺犪犻犪,2001,21(3): ( : )

BuletinofSoiland WaterConservation Vol.38,No.3 Jun.,2018 & X R( + 1, 1, < 1,=> 3 (1. E^ IFTU ^,NO ; 2.NO\ ^,NO ;3.NO B`a -

$BuletinofSoiland WaterConservation Vol.38,No.3 Jun.,2018 & X R( + 1, 1, < 1,=> 3 (1. E^ IFTU ^,NO ; 2.NO\ ^,NO ;3.NO B`a -$ 383 0186 BuletinofSoiland WaterConservation Vol.38,No.3 Jun.,018 & X R( + 1, 1, < 1,=>,?@A 3 (1. E^ IFTU ^,NO 730070;.NO\ ^,NO 730070;3.NO B`a -[/H,NO 730000) -.:[# ] _D5 & % I /, + -. I P+- [ ] ( 犜狉犻狋犻犮狌犿犪犲狊狋犻狏狌犿