325 201810 JournalofSoiland WaterConservation Vol.32No.5 Oct.,2018 KZ[%@,;3@Y\]^ "_`ap O 1,<) 2,3,4,*73 2,3,4,+ 1,2,3,4,E9 5 (1.?Q345;,? 712100; 2.;<6)*6O"0, 100101; 3.;<)*Q6"0,? 712100; 4.;3645;, 100049;5.?%&`,? 719399) :.:NA0,)*+JQ_`( )@ 6 F H(F 犘狀 b 犌狊 8 CO2 犆犻 <[ WUE) 犜狉 MJ Ψ 狑 V \:+_`@ 犘狀 犌狊 WUE M 犜狉 c V, 犆犻 Ψ 狑 U ]a @ J V 2,0`+:, 犘狀 犌狊 ]a, @ 犘狀 犌狊 ] a ( 狆 <0.05)(+C) +,@ WUE _ `8D]a_^( 狆 <0.05), @ 0,Q )6+:_Z', Z = > _ ` J +V_`Z :+;;;@ ;<Z ()*+,:S728.2 -./0:A -1,:10092242(2018)05023406 犇犗犐 :10.13870/j.cnki.stbcxb.2018.05.038 犈犳犲犮狋狊狅犳犇狉狅狌犵犺狋犛狋狉犲狊狊狅狀狋犺犲犘犺狔狊犻狅犾狅犵犻犮犪犾犪狀犱犈犮狅犾狅犵犻犮犪犾犆犺犪狉犪犮狋犲狉犻狊狋犻犮狊狅犳 犃犿狔犵犱犪犾狌狊狆犲犱狌狀犮狌犾犪狋犪犘犪犾犪狀犱犛犪犾犻狓狆狊犪犿犿狅狆犺犻犾犪犛犲犲犱犾犻狀犵狊犻狀犛狅犻犾狊狑犻狋犺犇犻犳犳犲狉犲狀狋犜犲狓狋狌狉犲狅狀狋犺犲犔狅犲狊狊犘犾犪狋犲犪狌 PEIYanwu 1,HUANGLaiming 2,3,4,JIA Xiaoxu 2,3,4,SHAO Ming an 1,2,3,4,ZHANG Yinglong 5 (1. 犆狅犾犲犵犲狅犳犖犪狋狌狉犪犾犚犲狊狅狌狉犮犲狊犪狀犱犈狀狏犻狉狅狀犿犲狀狋, 犖狅狉狋犺狑犲狊狋犃牔犉犝狀犻狏犲狉狊犻狋狔, 犢犪狀犵犾犻狀犵, 犛犺犪犪狀狓犻 712100; 2. 犓犲狔犔犪犫狅狉犪狋狅狉狔狅犳犈犮狅狊狔狊狋犲犿犖犲狋狑狅狉犽犗犫狊犲狉狏犪狋犻狅狀犪狀犱犕狅犱犲犾犻狀犵, 犐狀狊狋犻狋狌狋犲狅犳犌犲狅犵狉犪狆犺犻犮犛犮犻犲狀犮犲狊犪狀犱犖犪狋狌狉犪犾犚犲狊狅狌狉犮犲狊犚犲狊犲犪狉犮犺, 犆犺犻狀犲狊犲犃犮犪犱犲犿狔狅犳犛犮犻犲狀犮犲狊, 犅犲犻犼犻狀犵 100101;3. 犛狋犪狋犲犓犲狔犔犪犫狅狉犪狋狅狉狔狅犳犛狅犻犾犈狉狅狊犻狅狀犪狀犱犇狉狔犾犪狀犱犉犪狉犿犻狀犵狅狀狋犺犲犔狅犲狊犘犾犪狋犲犪狌, 犐狀狊狋犻狋狌狋犲狅犳犛狅犻犾犪狀犱犠犪狋犲狉犆狅狀狊犲狉狏犪狋犻狅狀, 犆犃犛牔犕犠犚, 犢犪狀犵犾犻狀犵, 犛犺犪犪狀狓犻 712100;4. 犆狅犾犲犵犲狅犳犚犲狊狅狌狉犮犲狊犪狀犱犈狀狏犻狉狅狀犿犲狀狋, 犝狀犻狏犲狉狊犻狋狔狅犳犆犺犻狀犲狊犲犃犮犪犱犲犿狔狅犳犛犮犻犲狀犮犲狊, 犅犲犻犼犻狀犵 100049;5. 犛犺犲狀犿狌犈犮狅犾狅犵犻犮犪犾犃狊狅犮犻犪狋犻狅狀, 犛犺犲狀犿狌, 犛犺犪犪狀狓犻 719399) 犃犫狊狋狉犪犮狋 :Thepresentstudyinvestigatedthephotosyntheticcharacteristics (netphotosyntheticrate ( 犘狀 ), stomatalconductance ( 犌狊 ),intercelularco 2 concentration ( 犆犻 ),internalwateruseeficiency (WUE)), transpirationrate( 犜狉 )andleafwaterpotential(ψ 狑 )of 犃犿狔犵犱犪犾狌狊狆犲犱狌狀犮狌犾犪狋犪 Paland 犛犪犾犻狓狆狊犪犿犿狅狆犺犻 犾犪 underdroughtstressinsoilswithdiferenttextures (sandysoilandloamysoil)ontheloessplateau throughwatercontrolpotexperiment.theresultsshowedthat 犘狀, 犌狊,WUEand 犜狉 of 犃. 狆犲犱狌狀犮狌犾犪狋犪 Pal and 犛. 狆狊犪犿犿狅狆犺犻犾犪 insoilswithdiferenttexturesincreasedfirstlyandthendeclinedunderthedrought stress,while 犆犻 and Ψ 狑 showedatrendofrisinganddecreasing,respectively.soiltexturesignificantly afectedthesensitivityof 犃. 狆犲犱狌狀犮狌犾犪狋犪 Paland 犛. 狆狊犪犿犿狅狆犺犻犾犪 tothewaterdeficiency.underthesame droughtdegree,the 犘狀 and 犌狊 of 犃. 狆犲犱狌狀犮狌犾犪狋犪 Palintheloamysoilweresignificantlyhigherthanthosein thesandysoil,whileanoppositetrendwasobservedforthe 犛. 狆狊犪犿犿狅狆犺犻犾犪 ( 狆 <0.05)(exceptthesevere :20180409 :!"(41601221); ; ) * Q 6 " 0! " (A3140214021602);;<6)*<!"(2017RC203);6O"0 <)**!"(LENOM2016Q0001) :(1992 ),,()*,,-./<)* Email:peiyanwuxn@163.com <=:\7(1956 ),,)*+,,-./<)* Email:shaoma@igsnrr.ac.cn QA\(1984 ),,)*+,,-./<6N)* Email:huanglm@igsnrr.ac.cn
5 :+JQ_`@ 6<HIV 235 droughttreatment).asignificantdiferencein WUEof 犃. 狆犲犱狌狀犮狌犾犪狋犪 Paland 犛. 狆狊犪犿犿狅狆犺犻犾犪 wasob servedbetweenthesoilswithdiferenttextureswhenthesoilmoisturewasreducedtomoderateandsevere drought( 狆 < 0.05).Thegrowthof 犃. 狆犲犱狌狀犮狌犾犪狋犪 Palintheloamysoilwasbeterthanthatinthesandy soil,while 犛. 狆狊犪犿犿狅狆犺犻犾犪 showedanoppositetrend.therefore,duringthevegetationrestorationandecologicalcon structionontheloessplateau,thetreespeciesshouldbechosennotonlyaccordingtothesoilwatercondition,butalso consideringthediferentresponsesofvegetationtodroughtstressinsoilswithdiferenttextures. 犓犲狔狑狅狉犱狊 :droughtstress;soiltexture; 犃犿狔犵犱犪犾狌狊狆犲犱狌狀犮狌犾犪狋犪 Pal; 犛犪犾犻狓狆狊犪犿犿狅狆犺犻犾犪 ;physiological response Q!0,S? V!,)6+S$Q -.U)V-#,AQ +J_U _< -.U) HUS 3,67V`[,bAQ _`!(?,+H [12] abv)* [3] \,Q)/b c!b*vff P +SQN\+JV- 28b [4] _78!1 4_I [5],\]^Q+? LG [` V!++HQ Y+V-JK,)*J +VZHIMZKJQ )6+D- D)* [68] \,_`M`W^ X<:J+V2_` abw J+Q4VH [6] 4CVJHI [8] P) *,\,4V)* a,de1,dv )*a_;4cqcvj HI+Va3,CC #,Ca,3(C C C)V+ Z]aQC,3)* [910] \, J+VZOZK: 8c(\]VH,G_` ^ZVBFF_` "+ Q@ V <HI DCc [68],WJc!@ @ _`VM)*`,M)*[.:NA0,Q @ )*J?,)*+J @ F H M J V,5L_`@ J+ ZV_^,Q+>+! )6+$%&' 1 +, 1.1 C M)*;<)* 645A0P,A0?% 01(110 21 110 23 E,38 46 38 51 N),= 1094~1273m c01@:k, VR@QZ[,S5 7 c01g6.9km 2," Q,b>+3b b 8.4,437mm,`3, 6 9V8V77%,GcO ( 013=>V,$ 45VW,c!1 #,,- @ XU D [12] @ S 9 V $@ [1112],bA :<!P3G_,+ > [13] 1.2 JK'P M)*MVW6 V@ 6,[.:A0V 645A0P=M,=<[Y O $ V + :. : [ c 30 cm,-25cm [01,M<@1 X+ (.2mm),+c6: 6 8 W, U 6 : 7,8kg 6:^.,V2=, WV@ L.L63Q 6:.,3.:A0 4 O( @ @ ), O5 ^.3QR2c(. 0.5~1.0cm),K 31X (A8)+A0,.: 80% FC(J I) U 60% FC,40% FC 20% FC,U@ 6FH(F 犘狀 b 犌狊 8 CO 2 犆犻 <[ WUE)M J
236 32 (Ψ 狑 )V,<[(WUE),):, 2=[ +=V 1 @2P / (g cm -3 ) / (cm h -1 ) 8/ /% R Da/ 8/ 8/ 1.65 13.44 147.72 5.15 31.28 63.58 2.07 1.32 1.38 1.36 8.23 274.35 13.88 51.13 34.99 3.49 1.54 1.67 :10 UVVL ` 1.3 ' [O CL 340FJ PSYPRO UJ^<FHP 8=(8:00 19:00),LL1/h ^ < ), 8 [ 2 = (14)L A 08bFIF@51 犆犻 _ ` + (60% FC,40% FC 20% FC)c _ ` 8 犆犻 ]a _ ^, @ 犆犻 + (40% FC)]a( 狆 <0.05) )1 M<bc( 犘犃犚 )UV 1.4 * [SPSS16.0VP',[\ (LSD)<8V_^]a(α=0.05), [ Origin9.0VP5 2 &-./ 2.1 KZ[;3@Y\]^"_`a M 52Y,+]a_` @ 6F( 犘狀 )b( 犌狊 ) ( 狆 <0.05) 6JI(CK 80% FC)01,2+ V _ ` @ 6 犘狀 犌狊 cv,\+d <@ V 0`+:(60% FC,40% FC,20% FC) 犘狀 犌狊 ]a,u(3.87,2.64,1.01μmol/(m 2 s)24.72,20.47,13.12 mmol/(m 2 s) 0, @ 犘狀 犌狊 + (60% FC) + (40% FC) ] a, U (3.21, 2.28μmol/(m 2 s) 18.79,15.67 mmol/(m 2 s),+(20% FC)_`@ 犘狀 犌狊 =D] a _ ^ _ ` @ 8 CO 2 ( 犆犻 )J + V Z H I _ ` 6JI (CK 80% FC)0 1, @ )2 ;@3Y\]^"_`aM UR CO2 6 2.2 KZ[;3@Y\]^"_`a 1?cW?*< 2Y,+]a_` @ J(Ψ 狑 )( 狆 <0.05)M <[ (WUE)( 狆 <0.01) 6 J I (CK 80% FC)01,60% FC <_ `J@ Ψ 狑 V`( 狆 > 0.05),W]a WUE( 狆 <0.01) 2 V,_`@ Ψ 狑
5 :+JQ_`@ 6<HIV 237 M WUE 6JI(CK 80% FC)0 1 D ] a _ ^,\ S @ Ψ 狑 M WUE V - + (40% FC),_ ` 8 @ Ψ 狑 M WUE D]a_^( 狆 <0.05),+ (20% FC)@ WUE ]a, \S@ Ψ 狑 M WUE V-,G_`_` 53Y,@ Ψ 狑 _` D0`V,8=`3,a3F a F U ( 10:00 14:00 6 J I (CK 80% FC)01,60% FC <J @ Ψ 狑 ` 3 40% FC,_`@ Ψ 狑,@ _`8 Ψ 狑 W" (40% FC 20% FC) ] a @ Ψ 狑 _ ` @ WUE 0`,c\,W_`<UV@ F( 8D_^ 6JI(CK 80% FC)01,+ <@ 6 WUE F c 60% FC U V @ _ ` 8 WUE, 40% FC UV@ _`8 WUE +(20% FC),_`@ Ψ 狑 WUE,G 2 @ @ ;3?*PY\]^ "_1?cW?*< < J /MPa <[/ ( μ mol mmol -1 ) CK(80% FC) -2.62±0.68 Aa 3.20±1.12 Aa 60% FC -2.84±0.43 Aa 4.84±0.95 Ab 40% FC -3.07±0.29 Ab 4.59±1.23 Ab 20% FC -3.39±0.18 Ac 3.76±0.78 Ac CK(80% FC) -2.68±0.25 Aa 3.08±1.01 Aa 60% FC -2.72±0.28 Aa 4.58±0.96 Ab 40% FC -2.89±0.11 Ab 4.82±1.04 Ab 20% FC --3.47±0.45 Ac 3.89±1.23 Ac CK(80% FC) -2.36±0.43 Ba 3.33±1.11 Aa 60% FC -2.52±0.21 Ba 4.03±1.08 Bb 40% FC -3.01±0.13 Ab 5.30±1.02 Bc 20% FC -3.33±0.26 Ac 3.75±1.13 Abd CK(80% FC) -2.16±0.23 Ba 2.60±1.08 Aa 60% FC -2.28±0.38 Ba 4.51±0.98 Ab 40% FC -2.80±0.16 Ab 5.01±1.15 Bc 20% FC -3.28±0.26 Ac 3.26±1.23 Bd :FC8;_`3"`0` _`8_^]a( 狆 <0.05);_`"` 0`_`+:8_^]a( 狆 <0.05) )3 ;3?*PY\]^"_1?cW?*<
238 32 2.3 KZ[;3@Y\]^"_`a +,YJV AV*8,ZSZJ+5 VWO 54Y,_` @ += c V + :(1~7=),`%&,_` @ 犜狉 ( N V +10=,,_ ` @ 犜狉 ( + 20 =, %,_` @ 犜狉 X+8, 犜狉 F` (7=,@ 犜狉 F`@ (4=,\ +@ 犜狉 V VFF [3],[`V+ Q)+V-JK M )*.:NA0(_` @ J+VZHI,JQ KP+D- F[SV,F@ SZ+V-HI#W [14] M)* \,+@ F@VZHI 6JI(CK 80% FC)01,_ `@ 犘狀 M 犌狊 + (60% FC) D, + (40% FC20% FC)]a 0`+:, 犘狀 犌狊 ]a,@ 犘狀 犌狊 ]a(+20% FC C ) \_`V _`, J + V Z, @ J+VZ F[V bb,s+ b,co 2 :JF, SVF Farquha [15] 犘狀 犌狊 犆犻,Sb, 犘狀 犌狊 犆犻,\Sb M)* _`@ 犘狀 M 犌狊 犆犻 D,F[,-bNK [16] J_`:+@JVF[ P)*,(0`V [17] )*\, bs+qu :F@V, )4 ;3@Y\]^"_`a @?HK 3 0 ' +SV-#W Q)/bc!b*,W]45UV(3b)J 犘狀 犌狊 D]a +45,Y<`3a` )#8V,.3J <[,ZSV- [9] M )*\,2+,_` @ WUE cv,. + ) $ @ WUE J @ [16] [18] [19] _`V)*D V Zhang [20] Zhang [21] J ` `F[ WUE V # FF FF,W=> J WUE V M)*\, @ WUE_, V +(40% FC), _` 8 @ Ψ 狑 M WUE D] a _ ^ ( 狆 <0.05), + (20% FC)@ WUE ]a,\ S @ Ψ 狑 M WUE V -,G_`_`,W\ S@ WUE V-
5 :+JQ_`@ 6<HIV 239,+, FF Z=>_` WUE Y)V]a_^ S)*V-@, YbA3,.(+ J < V [22] _ U,b 8,. 犜狉,,: J V CO 2, 犘狀 W +UV,bJV3 J CO 2 *VJK 6Fa` J2V_^./+ <[V$,ZV< [9] M)*\,_`@ 犜狉 +=c V,75g/kg (30% FC), 犜狉 犜狉 F`(7=,@ 犜狉 F`@ ( 4 =, \ +@ 犜狉 V 4 & ' +_`@ 6 F ( 犘狀 ) b ( 犌狊 ) < [ (WUE)M ( 犜狉 ) c V,8 CO 2 ( 犆犻 )(Ψ 狑 )U ]a@ JV2,0`+:, 犘狀 犌狊 ]a,@ 犘狀 犌狊 ]a( 狆 <0.05)(+C) +,@ WUE_` 8 D ] a _ ^ ( 狆 <0.05),QP)6+,_ Z',Z=> _`J+V_`Z -.: [1] \7,T,,.Q + ) * 6[J].,2016,31(1):1422. [2] Shao M A,WangY Q,XiaY Q,etal.Soildrought andwatercarryingcapacityforvegetationinthecritical zoneoftheloessplateau:areview [J].VadoseZone Journal,2018,17:e170077. [3] FengX M,FuBJ,PiaoSL,etal.RevegetationinChina s LoessPlateauisapproachingsustainablewaterresourcelimits [J].NatureClimateChange,2016,6(11):10191022. [4] DaiA.Erratum:Droughtunderglobalwarming:Are view [J]. Wiley Interdisciplinary Reviews Climate Change,2011,2(1):4565. [5] Liang W,BaiD,WangFY,etal.Quantifyingtheim pactsofclimatechangeand ecologicalrestoration on streamflow changes based on a Budyko hydrological modelinchina sloessplateau [J].WaterResources Research,2015,51(8):65006519. [6],U,>.+Q4 H[J].,2012,32(4):11741184. [7] XR,U, V,.Q > CJ + V < Z[J]. 645,2013,22(1):4349. [8],>, ),.+ J Q 4 C H I V [J].,2014, 34(16):45354548. [9] KatiyaragarwalS.Methodsandconceptsinquantifying resistancetodrought,saltandfreezing,abioticstresses thatafectplantwaterstatus[j].plantjournal,2006, 45(4):523539. [10] 7,>.ZJ+V[J]. Z[,2012,23(10):29072915. [11] JiaH X,YangH F,SunP,etal.Denovotranscriptome assembly,developmentofestssrmarkersandpopulation geneticanalysesforthedesertbiomasswilow,salixpsam mophila[j].scientificreports,2016,6:e39591. [12],\7,Q A \. 0 1 K a HIM[J].,2017,31 (5):222230. [13] Z,,`,.@@V Z[, CN200910021255.5 [P].20090722. [14] =,,,.+FFZ :O61`[J].,2017,37(7):23152324. [15] Farquhar G D,Sharkey T D.Stomatalconductance andphotosynthesis[j].annualreviewofplantphys iology,1982,33(3):317345. [16],F,,.+J@JF [V[J].,2013,33 (5):13861396. [17],[,,.+ Q U : V F +FH[J].Z[,2011,22(6): 13951402. [18] #,F,T,.JF<@J F I V F F Z [J]., 2011,35(3):322329. [19] N,,Q,._`JFH M<[ J V Z [J]., 2015,35(9):29022909. [20] ZhangGC,XiaJB,ShaoH B,etal.Gradingwood landsoilwaterproductivityandsoilbioavailabilityin thesemiaridloessplateauofchina[j].actahydro chimicaethydrobiologica,2012,40(2):148153. [21] ZhangS Y,Zhang G C,GuS Y,etal.Criticalre sponsesofphotosyntheticeficiencyofgoldspurapple treetosoilwatervariationinsemiaridloesshilyarea [J].Photosynthetica,2010,48(4):589595. [22] Cochard H.Anoverviewofmodelsofstomatalcon ductanceattheleaflevel[j].plantcel & Environ ment,2010,33(9):14191438.