Quercus aquifolioides Rehd. et Wils. Picea likiangensis var. linzhiensis Cheng et L. K. Fu Desmodium elegans DC. Leptodermis potaninii var. 7 g

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1 Forest Research * DCCA Quercus aquifolioides 1 2 > > DCCA 48 4 DCCA S A Pattern of Plant Species Diversity of Quercus aquifolioides Community in Southeast Tibet China WANG Guo-yan 1 2 LUO Jian 2 XU A-sheng 2 LU Chuang 1 1. Tibet Agricultural and Animal Husbandry College Nyingchi Tibet China 2. Research Institute of Tibet Plateau Ecology Nyingchi Tibet China Abstract The special topographic and climate conditions of Himalaya-Hengduan Mountains gestate a rich forest resource and plant and animal diversity in southeast Tibet. Through plot investigation the species diversity and spatial distribution of Quercus aquifolioides community in southeast Tibet were studied using diversity index and detrended canonical correspondence analysis DCCA combining with specific association interspecific covariation and systematic cluster analysis. The results indicated that 1 The species richness decreased with the raise of altitude but a certain degree of human disturbance i. e. moderate disturbance higher canopy density and lower canopy layer usually increased the species diversity. In addition the pattern of species diversity was influenced by the microclimate of dry valley. 2 The species diversities of different layers in the community appeared as herb layer > shrub layer > tree layer. 3 Theoretically the higher ratios of positive and negative association implied the more stable community structure and the ecological habits and distributional differences among species of Q. aquifolioides community formed different interspecific relationship. Moreover the distribution of species was mainly affected by altitudinal gradient and annual precipitation. 4 According to the outcomes of DCCA specific association interspecific covariation and systematic cluster analysis the main populations of Q. aquifolioides community could be divided into four ecological species groups it helps to understanding the local community structure and plant ecological habits and further studying on the community succession conservation and utilization. Key words Southeast Tibet Quercus aquifolioides species diversity DCCA * luojian-sh@ sohu. com

2 Quercus aquifolioides Rehd. et Wils. Picea likiangensis var. linzhiensis Cheng et L. K. Fu Desmodium elegans DC. Leptodermis potaninii var. 7 glauca Diels H. Winkl. L. purdomii 7-8 Hutch. Berberis hemsleyana Ahrendt 9-11 Populus davidiana Dode Rosa sericea Lindl. Cotoneaster microphyllus Wall. & Lindl. var. cochleatus Franch. Rehd. & 14 Wils. Lonicera inconspicua Batal. Potentilla fruticosa L ' 30 00' N 94 19' 94 53' E georgei var. smithii Cheng et L. K. Fu Drynaria sinica Diels Thalictrum alpinum L. Leibnitzia nepalensis Kunze Kitam. Potentilla Griffithii Hook. f. Salvia roborowskii Maxim m Triplostegia glandulifera Wall. ex DC Stipa capillacea Keng Chamaesium paradoxum Wolff Aster souliei Franch. Orinus thoroldii Stapf Bor Smilax menispermoidea A. DC. Clematis montana Buch. -Ham. et DC mm 64% cm 15 Pinus densata Mast m Betula utilis D. Don 16 P. armandi Franch. Hippophae rhamnoides L. Abies 20 m m 2 20 m 1 9

3 Rhododendron wardii W. W. Smith Berberis johannis Ahrendt Lonicera microphylla Willd. ex Roem. et Schult. Elsholtzia fruticosa D. Don Rehd. Cotoneaster multiflorus Bge. Ribes glaciale Wall. Rosa macrophylla Lindl var. glandulifera Yü et Ku Lonicera webbiana Wall. & DC. Ribes himalense Royle ex Decne. Lonicera lanceolata Wall. Sorbus rehderiana Koehne Rosa omeiensis Rolfe Viola szetchwanensis W Beck. et H. de Boiss. var. nudicaulis W. Beck. Ligularia myriocephala Ling ex S. W. Liu Polygonum nepalense Meisn. Ligularia tsangchanensis Franch. Hand. -Mazz. Circaea alpine Linn. Viola szetschwanensis W. Beck. et H. de Boiss. Geranium wilfordii Maxim. Poa chalarantha Keng ex L. Liu Halenia elliptica D. Don var. elliptica Elymus nutans Griseb. Dryopteris silaensis Ching Pteris dactylina Hook /m 1 / / /mm 4 /m Q Q Q Q Q Q Q Q Q Q Q Q Q

4 Q Q1 Q4 Q6 Q12 Simpson /Shannon-Win- ner / / / / m 2 m 1 m 1 m 2. 2 = + + /3 = + + /3 Simpson D Shannon-Winner H Pielou J DCCA Detrended Canonical Correspondence Analysis Q5 Q11 Simpson /Shannon- 2 2 Visual Foxpro 6. 0 Winner 0. 8 / /2. 24 Yates X 2 Q Q11 SPSS Spearman Q Simpson DPS Shannon-Winner 3 2 CANOCO DC- CA 群落层次间多样性的比较 不同样地物种多样性的比较 Simpson Shannon-Winner Compositae Rosaceae Simpson D Shannon-Winner H Pielou J Q10 Q2 Q2 2 > > Q10

5 6 707 Q2 3 4

6 X X Spearman Spearman > 34-80% 35 5 X 2 Spearman DCCA

7 6 709 DCCA 33 7 Ele Cand Slop Posi Soiw Anpr Asp Antd 7 - DCCA > > DCCA DCCA Q Simpson Simpson Simpson

8 DCCA DCCA A m B m P < C m D X DCCA 4. 3

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