32 o 30 42 o 30 103 o 30 124 o 10 1980 1500m 3000m 8 14 3 1-13 -2-30 25 28 40 150 230 600 900mm 3 7 10 40 60 70 10 4 5 9 4.1 Chen 1995
1981 1987 1981 1985 1992 1986 1988 1990 1992 1994 Chen 1995 1,5 7 11 2,3 8,12 13 4 9 14-18 6 10 19-22 4.2 TWINSPAN 92 92 22 4 2 Vitex negundo var heterophylla Artemisia gmelinii + Carexreventa D 1 D 2 D 4 D 5 D 4 D 5 Galium aparine Cacalia hastata + Larix principis-rupprechtii Picea meyeri Picea wilsonii Pinus armandii
D 2 55 12 1 5 1800 2500m 6 12 Betula chinensis 1300 1800m D 3 Platycladus orientalis + Pinus bungeana D 6 D 7 Arundinella hirta Bidens parviflora + D 3 37 10 13 14 1050 1200m 1500 1650m 15 18 200 800m 19 20 370 400m 21 1050m 22 250 350m TWINSPAN 4 2 DCA 103 403 DCA 4.3 4.4 103 X 1 X 2 X 1 X 3 X 4 K X 1 93 169 X 3 26 286 X 4 8 299 X 3 X 4 I X 1 360 564 X 2 238 368 X 3 118 177 X 4 100 235 X 1 X 1 X 2 X 4 A E D E X 1 B C X 1 X 2 X 3 X 4 0 261 A E X 1 F X 1 218 276 X 2 48 180 X 3 155 299 X 4 5 156 X 2 X 3 X 4 A G X 1 X 2 X 3 X 4 H N L H N M L X 1 X 2 X 3 H X 1 49 121 X 2 172 241 X 3 101 193 X 4 90 227 X 1 56 113 X 2 200 214 X 3 92 116 X 4 182 205 L M X 1 7 61 0 49 X 2 190 221 207 218 X 3 107 169 145 165 X 4 172 194 126 158 X 1 X 1 X 1
4.3 X 1 X 2 DCA
4 4 X 1 X 3 DCA (K) (D E) (I) 1000m
A B C F 1300 1800m G H L M N 2500m 4 3 Rhamnus parvifolia Prunus armeniaca Myripnois dioica Lespedeza dahurica 4 1 Carex humilis var nana Cleistogenes spp Deyeuxia arundinacea Atractylodes chinensis Lespedeza hedysariodes Rhus chinensis Carex prevernalis C humilis var nana C rigescens Platycodon grandiflorum Artemisia japonica var mandshurica Cynanchum acuminatifolium Oxalis acetosella 4.1 Fraxinus rhynchophylla 2 Pinus tabulaeformis 5 Platycladus orientalis 5 Quercus dentata 2 Q.liaotumgensis 4 Berberis poirelii 2 Campylotropis macrocarpa 2 Fraxinus bungeana 2 Indigofera bungeana 2 Leptodermis oblonga 2 Lespedeza bicolor 3 L.dahurica 2 L.floribunda 5
L.hedysaroides var.inschanica 3 Leptopus chinensis 2 2 Myripnois dioica 2 2 Prunus davidiana 2 P.armeniaca var.ansu 3 Rhamnus globosa 2 R.parvifolia 2 Rhododendron micranthum 2 Spiraea pubescens 4 S.trilobata 4 Vitex negundo var.heterophylla 4 5 Artemisia gmelinii 5 4 Arundinella hirta 3 2 Atractylodes chinensis 4 Bupleurum falcatum 2 B.pekinesis 3 2 Carex humilis var.nana 3 3 C.lanceolata 2 Cleistogenes serotina 2 3 C.squarrosa 2 Dendranthema zawadskii 3 Deyeuxia arundinacea 2 Iris dichotoma 2 I.ruthenica 2 Leibnitzia anandria 3 Leontopodium leontopodioides 2 Polygala sibirica 2 P.tenuifolia 3 Polygonatum macropodium 2 Potentilla chinensis 2 Pulsatilla chinensis 2 Rhaponticum uniflorum 2 Sanguisorba officinalis 3 Saposhnikovia divaricata 2 Scorzonera glabra 2 Selaginella sinensis 3 Spodiopogon sibiricus 3 4 2 Rhododendron micranthum Phlomis umbrosa Rodgersia aesculifolia Carpinus turczaninowii Abelia bif-
lora Spiraea trilobata Viburnum betulifolium Rosa bella Acer hersii 2 A.mono 2 Betula chinensis 2 Carpinus cordata 4 C.turczaninowii 2 1 Crataegus pinnatifida 2 3 Malus baccata 2 Pinus armandii 2 5 P.tabulaeformis 5 3 Quercus aliena var.acuteserrata 3 Q.liaotungensis 4 Rhus potaninii 2 R.verniciflua 2 2 Tilia henryana var.subglabra 2 T.mandshurica 2 Abelia biflora 2 2 Berberis amurensis 2 Corylus heterophylla 2 2 C.mandshurica 2 Elaeagnus umbellata 2 2 Evonymus alatus 2 Forsythia suspensa 2 2 Lespedeza bicolor 4 2 L. Buergeri 2 Lonicera maximowiczii 2 Ostryopsis davidiana 2 Rhododendron micranthum 2 Rosa bella 4 3 R.xanthina 2 Rubus idaeus 2 Spiraea pubescens 4 S.trilobata 2 Viburnum betulifolium 2 2 Agrimonia pilosa 2 Agropyron mongolicum 2 Anemone vitifolia 3 2 Artemisia gmelinii 2 Aster ageratoides 4 4 Atractylodes chinensis 2 Bupleurum chinensis 2 Carex duriuscula 2 4
C.lanceolata 2 C.reventa 4 C.siderosticta 3 Carpersium cernuum 2 Cassia tora 2 Dendranthema chanetii 4 Deyeuxia arundiancea 2 5 Euphorbia lunulata 2 Iris ruthenica 3 Maianthemum bifolium 2 Patrinia heterophylla 3 Pedicularis striata 2 Phlomis umbrosa 3 Polygala sibirica 2 Polygonatum sibiricum 2 Pulsatilla chinensis 2 Rodgersia aesculifolia 2 Rubia cordifolia var.longifolia 3 Sanguisorba officinalis 2 Sanicula chinensis 2 Saposhnikovia divaricata 2 Sanssurea japonica 3 Smilax china 2 Vicia unijuga 2 Viola acuminata 2 Dendranthema chanetii Fragaria orientalis 4 3 Betula allbo-sinensis 3 1 2 B.platyphylla 3 4 Larix principis-rupprechtii 5 5 5 Picea meyeri 2 5 4 P.wilsonii 1 3 4 Salix heterochroma 2 Abelia biflora 1 2 Berberis amurensis 3 Cornus alba 2 3 Corylus mandshurica 3 3
Cotoneaster acutifolius 3 2 C.zubelii 2 1 Crutaegus kansuensis 2 Daphne giraldii 3 Evonymus alatus 2 3 Lespedeza bicolor 2 1 2 Lonicera caerulea var.edulis 2 L.chrysantha 3 3 4 L.hispida 2 3 2 L.maackii 4 Potentilla glabra 2 P.fruticosa 4 Ribes burejense 2 R.mandshuricum 2 2 Rosa bella 5 2 4 Rubus idaeus 2 3 Salix phylicifolia 2 Spiraea pubescens 5 5 Viburnum mongolica 2 2 Aconitum kusnezofii 2 1 1 Aegopodium alpestre 3 Agrimonia pilosa 4 1 Allium sacculiferum 3 Anaphalis hancockii 2 Aquilegia yabeana 3 Aster ageratoides 1 2 4 Bupleurum chinensis 4 2 2 Cacalia hastata 2 1 Cardamine tangutorum 2 2 Carex lanceolata 2 1 C.siderosticta 4 5 5 Carpesium cernuum 3 1 Cerastium arvense 2 1 Cicuta virosa 2 1 Cimicifuga dahurica 2 Clematis macropetala 3 Convallaria keiskei 2 Cortusa mathiolis var.pekinensis 4 Dendranthema chanetii 4 3 5 D.zawadskii 2 Deyeuxia arundinacea 2 1 Dianthus superbus 2 Duchesnea indica 2 1 Equisetum arvense 4 E.pratense 5 Fragaria orientalis 2 3 4
Galium aparine 2 4 G.bungei 2 5 G.verum 2 2 Gentiana macrophylla 1 2 Geranium sibiricum 3 2 Halenia corniculata 2 Herminium monorchis 3 Iris ruthenica 2 2 Lathyrus hunmilis 3 5 Maianthemum bifoiium 1 2 4 Melica radula 3 3 Melica scabrosa 3 4 3 Pedicularis muscicola 4 Phlomis umbrosa 2 4 Poa.sp 1 4 Polemonium coeruleum var.chinensis 3 1 4 Polygonatum odoratum 1 3 P.sibiricum 2 2 P.viviparum 1 4 1 Ranunculus brotherusii 3 4 2 Sanguisorba officinalis 1 2 Saposhnikovia divaricata 2 2 Thalictrum petaloideum 3 1 T.thunbergii 1 3 Veratrum nigrum 3 Vicia unijuga 3 2 4 Viola biflora 3 4.4 120 T-ANN 66.980150-0.282088 LONG - 0.578548 LAT - 0.005485 ALT T-JAN 68.624210-0.213677 LONG - 1.267432 LAT - 0.004826 ALT T-JUL 68.660480-0.334135 LONG - 0.081079 LAT - 0.006135 ALT PANN -2188.058000 + 42.251870 LONG - 56.545750 LAT + 0.137777 ALT EV-ANN 5465.375000-42.381790 LONG + 35.112920 LAT - 0.358505 ALT RH-ANN 22.574250 + 1.062319 LONG - 2.189187 LAT + 0.002190 ALT TSUM-10 16467.530000-76.378080 LONG - 86.410460 LAT - 1.605358 ALT BT 48.539990-0.197889 LONG - 0.332016 LAT - 0.004089 ALT K -58.370690 + 0.719988 LONG - 0.574378 LAT + 0.005721 ALT PER 8.185567-0.077859 LONG + 0.057020 LAT - 0.000497 ALT 103 DCA 4 4
3 9 9 4 1-12 4-5 8 7 18 25 7 483 786mm 7 K 3 8 10 7 0 7 1 3 6 0 7 5 7 5 5 9 3 7-2 4 3 4 1-16 -13 7 12 5 16 9 620 700mm 1340mm 11 14 1 7 10 A 111.98-34.53-1330.0-6.3- -7.7-18.7-753.9-1399.8-67- 2328.2-8.2-10.0-.7-112.02 35.45 1570.0 8.1-5.4 20.3 774.0 1455.1 69 2795.3 9.5-10.9.8 B 111.52-34.53-1420.0-3.9- -12.3-18.0-578.2-1412.1-60- 1919.8-6.8-8.8-.7-112.10 38.75 1650.0 7.6-5.8 19.7 786.4 1525.7 69 2650.8 9.1 10.7.9 C 111.52-36.62-1540.0-5.2- -10.4-18.7-604.6-1446.5-62- 2202.7-7.6-9.1-.8-112.05 37.72 1550.0 5.7-9.1 18.8 689.3 1507.8 65 2273.0 7.9 10.1.9 D 116.47-40.57-720.0-5.5- -12.4-21.0-525.0-1647.0 58-2602.3-8.2-6.2-1.0-116.65 41.02 880.0 6.6-11.3 22.0 554.2 1703.8 59 2890.6 9.0 7.1 1.1 E 116.30-40.37-210.0-6.9- -11.2-22.4-483.6-1720.0-58- 2985.3-9.2-3.8-1.1-117.13 40.92 650.0 9.4-8.6 25.0 551.9 1856.2 60 3681.9 11.0 5.9 1.3 F 111.93-33.72-1530.0-6.0- -6.6-17.5-845.6-1259.3-71- 2113.9-7.8-11.6.5 111.95 33.72 1800.0 7.5-5.3 19.1 883.5 1356.8 72 2548.6 8.9 13.2.6 G 111.50 37.83 1820.0-3.4- -12.1-16.9-634.5-1399.6-62- 1688.2-6.3-10.6-.7-1865.0 3.7-11.8 17.2 640.7 1415.8 62 1760.4 6.5 10.8.8 H 111.47-37.83-1865.0-.0- -15.8-13.7-609.9-1235.5-61- 728.6-3.9-10.7-.5-113.67 39.12 2350.0 3.4-12.1 16.9 701.4 1407.4 63 1688.2 6.3 13.5.7 I 111.83-36.47-1050.0-7.6- -7.6-10.9 614.1-1573.8-64- 2820.1-9.3-7.2-1.0-111.88 36.75 1200.0 8.6-6.5 21.9 642.5 1633.2 64 3085.4 10.0 8.1 1.0 K 116.47-40.33-270.0-5.5- -12.3-20.8-493.6-1622.5-58- 2566.8-8.2-3.9-1.0-117.12 40.85 910.0 9.3-8.6 24.8 565.0 1844.5 60 3645.7 11.0 7.5 1.3 L 111.47-37.87-2040.0- -.9- -16.6-12.5-661.5-1167.2-62- 474.9-3.1-11.8-.4-111.95 38.75 2250.0-2.4-12.9 15.8 702.2 1339.5 63 1406.9 5.6 14.6.6 M 111.95-38.75-2000.0-.3- -15.5-14.0-624.1-1256.8-61- 866.4-4.1-11.4-.5-112.12 38.92 2300.0 1.9-14.1 15.8 667.8 1363.1 62 1344.5 5.3 13.1.7 N 111.47-37.83-1960.0-1.6- -13.7-14.8-650.5-1282.1-62- 1150.1-4.9-11.3-.6-111.50 37.87 2200.0 2.9-12.5 16.3 683.5 1368.2 63 1535.3 5.9 12.7.7
4.5 TWINSPAN 166 559 TWINSPAN 4 5 Robinia pseudoaca- 4.5 TWINSPAN 1 7 13 2 8 14 3 9 15 4 10 16 5 11 17 6 12 18 cia + D 1 D 2 D 3 Acer hersii Astilbe chinensis + D 6 D 7 D 4
+ D 8 D 9 + Quercus aliena var. acuteserrata D 9 + D 10 D 14 D 14 11 + 12 Betula albo-sinensis + 166 18 (2) (3) (5) (4) (6) (7) (8) (Tilia mongolica) (9) (10) (11) 12 13 14 TWINSPAN DCA 1983 1984 1985 1989 1990 Chen 1995 4 6 DCA 166 559 DCA 4 6 P X 1 N X 1 442 458 X 2 143 175 X 3 98 179 X 4 80 140 DCA M X 1 326 483 X 2 115 285 X 3 87 223 X 4 80 350 L N X 1 287 434 K X 1 X 1 243 383 X 2 41 294 X 3 105 150 X 4 32 325 X 1 X 2 X 4 X 3 G H X 1 X 1 96 230 X 2 1 393 1 100 X 3 23 155 X 4 83 263 DCA 4 T X 1 179 253 X 2 273 357 X 3 152 306 X 4 141 259 (J) (G) X 1 141 212 X 2 35
4.6 X 1 X 3 DCA 94 X 3 68 97 X 4 170 203 A X 1 X 2 X 1 X 2 63 276 0 283 X 3 103 184 X 4 18 213 DCA C X 1
47 97 D F X 1 41 174 C X 3 145 267 X 1 0 51 X 3 C 101 184 X 1 X 3 X 1 Salix Caprea X 1 X 3 0 360 X 3 0 360 X 2 DCA 4.6 4 5 200 700m 800m Celtis koraiensis C bungeana Rhododendron mucronulatum Lindera obtusiloba Symplocos paniculata Zanthoxylum schinifolium Leptopus chinensis Lespedeza floribunda Polygonatum odoratum Cacalia aconitifolia Spodiopogon sibiricus Acer mono 2 2 3-2 A.truncatum 2 3-2 2 2 Albizia kalkora 3 Betula chinensis 3-2
B.costata 2 B.davurica 3-2 2 B.platyphylla 3-2 3 B.schinidtii 2 Carpinus cordata 2 1 C.turczaninowii 2 3-2 1 2 Castanea seguinii 1 Celtis koraiensis 2 2 2 1 2 2 C.bungeana 2 2 Fraxinus chinensis 2 F.mangshurica 2 2 F.rhychophylla 2 2 3-2 2 2 Juglans mandshurica 2 3 Juniperus rigida 2 5 Kalopanax septemlobus 2 Maackia amurense 2 2 2 Micromeles alnifolia 2 2 2 Phellodendron amurense 2 2 Pinus armandii 3 P.densiflora 2 2 2 P.tabulaeformis 2 2 2 Populus davidiana 3-2 2 3 Prunus armeniaca var.ansu 2-1 2-1 2 Quercus acutissima 3-5 2 Q.aliena 2 5 1 5 2 2 Q.aliena var.acutiserrata 3 5 Q.baronii 1 Q.dentata 2 2 5 2 3 Q.glandulifera var. 1 Brevipetiolata Q.liaotungensis 3-2 5-2 1 Q.mongolica 5 2 3-2 2 2 2 Q.variabilis 3 1 2 5 Rhus verniciflua 3 Tilia amurensis 2 T.henryana 1 T.mandshurica 2 2 3-2 Ulmus davidiana 2 U.laciniata 2 U.macrocarpa 3 U.propinqua 2 2 Abelia biflora 4-1 Acer griseum 2 Atractylodes japonica 2 Campylotropis macrocarpa 3 Celastrus articulata 2
Corylus heterophylla 4-1 4-1 3 2 3-1 C.mandshurica 4 2 Deutzia prunifolia 4-1 1 1 Evonymus sacrosancta 4-1 1 3 1 Exochorda serratifolia 3-1 Forsythia suspensa 3 Fraxinus bungeana 3 Grewia biloba var.parvifora 2 Indigofera kirilowii 2-1 4-1 2 1 2-1 Leptopus chinensis 2 1 1 Lespedeza bicolor 4-1 4-1 1 5 1 3 2-3 L.buergeri 3 L.davurica 2-1 1 1 1 3 L.floribunda 1 1 3 2 2 L.hedysaroides 3 var.inschanica L.formosa 3 Lindera obtusiloba 4 1 1 Lonicera praeflorus 2-1 1 1 Marlea macrophylla 2 Myripnois dioica 4 1 Neillia sinensis 3 Ostryopsis davidiana 2 3 Prunus padus 3 Philadelphus pekinensis 2 1 2 Rhamnus argutus 2 1 R.meyeri 2 R.parvifolium 2-1 1 1 R.schneideri var.manshurica 2-1 1 1 1 Rhododendron mirranthum 4-1 1 3 Rh.mucronulatum 4-1 1 Rhus potaninii 2 Rosa xanthina 2 Rubus cratagifolius 2-1 1 R.idaeus 2 R.saxatilis 2-1 Securinega suffruticosa 1 1 1 1 1 2 Sophora flavescens 3 Sorbaria sorbifolia 2 Spiraea frischiana 3 S.japonica var.fortunei 3 S.media 2-1 S.pubescens 4-1 4-1 3 S.trilobata 4-1 3-1 1 1 1 Styrax japonica 2 Symplocos panciculata 1 3 1 1 Syringa microphylla 4
S.velutina 2 Viburnum sargentii 2-1 1 1 Vitex negundo var.heterophylla 1 4 4 3 Weigela florida 2 Achnatherum pubicalyx 3-1 1 A.sibiricum 1 3 Adenophora tracheloides 2-1 1 1 2-1 Agrimonia gmelinii 2-1 1 1 Artemisia gmelinii 5 A.laciniata 3-2 A.stelonifera 2-1 Atrachylodes chinensis 2-1 1 1 A.japonica 2-1 1 1 Arthyrium crenatum 2 Arundinella hirta 2-1 3-1 Asparagus oligoclonos 2-1 1 1 Aster scaber 2-1 1 Athyrium crenatum 2 Bidens pubicalyx 2 Bothrichloa ischaemum 3 Cacalia aconitifolia 2-1 1 1 1 1 Carex callitrechos 3-2 C.campylorhina 1 3 C.humilis 3-2 3 3 C..humilis var.nana 3 C.lanceolata 3-1 3 3 3 3 3 3 C.nanella 3-2 4 2 3 3 2 C.quadriflora 3-2 C.siderosticta 3-1 3-2 4 C.ussuriensis 3-2 Cleistogenes caespitosa 2-1 3-1 1 2 C.hancei 2 Convallaria keiskei 2-1 1 2 Dendranthema lavandulaefolium 2 1 2-1 D.zawadskii 2 Deyeuxia arundinacea 2 3 5 D.zawadskii 3 Diarrhena yabeana 3 Dictamns dasycarpus 2 Dioscorea nipponica 2-1 1 1 2-1 Filifoliun sibiricum 3 Fragaria orientalis 2 Geranium dahuricum 2 Isodon glaucocalyx 3-1 Iris uniflora var.koreana 2-1
Melampyrum roseum 2-1 3 1 3 2 Meehania urticifolia 3 Phlomis umbrosa 3 Platycodon grandiflorum 2-1 1 1 1 2-1 Polygonatum odoratum 3-1 3-1 1 var.pluriflorum Potentilla chinensis 3 P.fragarioides 2-1 1 1 1 Pteridium aquilinum var.latiusculum 2 Pyrola dahurica 2 Sanguisorba parviflora 3-1 1 1 1 Saussurea mongolica 3 Scutellaria baicalensis 2 1 Sedum aizoon 2-1 1 1 Smilax stans 4 Spodiopogon sibiricus 4-1 3-1 3 3-2 Thalictrum baicalense 3 T.contortum 2 T.ichangense 3-2 Themeda triandra var.japonica 1 1 1 2 Vicia cracca 3-2 V.unijuga 2-1 3 1 80 600m 1000 1700m 1500 2200m 1982 Ulmus davidiana Rhus verniciflua Acer ginnala Evonymus sacrosancta Alangium chinense Meehania urticifolia Atractylodes japonica Lespedza formosa Forsythia suspensa Styrax japonica
Securinega suffruticosa Dioscorea nip ponica Themeda triandra var japonica Melampyrum Potentilla fragarioides Syringa villosa Lonicera tatarinovii Berberis amurensis Pyrola rotundifolia Trientalis europaea Gymnocarpiumcontinentalis Cardamine leucantha 4 6 Viburnum mungolicum Veratrum nigrum Saussurea nivea Thalictrum thunbergii Dendranthema zawadskii Acer mono 3 2 2 4 Betula platyphylla 5 2 1 2 B.albo-sinensis 2 5 1 B.chinensis 2 B.dahurica 3 2 5 2 Fraxinus rhynchophylla 2 Larix principis-rupprechtii 2 Populus davidiana 4 2 2 5 Quercus liaotungensis 3 3 Salix caprea 3 3 2 Sorbus pohuashanensis 2 2 1 Tilia mandshurica 2 T.mongolica 2
Abelia biflora 5 2 2 Eleutherococcus senticosus 1 Berberis amurensis 1 Cornus alba 3 C.bretschneidera 2 Corylus heterophylla 4 C.mandshurica 5 1 Lespedeza bicolor 3 1 4 Lonicera chrysantha 2 1 L.tatarinovii 2 Rosa xanthina 1 Spiraea trilobata 4 S.pubescens 5 2 4 Syringa pekinensis 2 S.villosa 2 Viburnum mongolicum 3 Agrimonia pilosa 2 Atractylodes chinensis 3 Aquilegia yabeana 2 Aster ageratoides 2 Cardamine leucantha 2 Carex lanceolata 4 3 3 5 C.siderosticta 2 3 Clematis macropetala 3 Convallaria keiskei 3 1 Dendranthema zawadskii 2 Gymnocarpium continentalis 3 2 Maianthemum bifolium 2 3 1 Phlomis umbrosa 4 Pyrola rotundifolia 2 Sanguisorba officinalis 2 Saussurea nivea 4 Spodiopogon sibiricus 3 Vicia pseudo-orobus 1 V.unijuga 4 Veratrum nigrum 3 Thalictrum thunbergii 3 Trientalis europaea 2 4.7 8 9 9 5 7 3 11
1-9 1-5 4-9 9-2 3 7 20 24 657 672mm 500 780mm K DCA X 1 X 2 X 3 1 7 10 PER K 5 4 9 0 1-7 4-4 7 K 1 7 10 A 116.75-36.33-300.0-8.9- -9.1-22.7-508.8-1561.7-59- 3375.6-10.7-4.4-.8-117.05 40.50 645.0 9.5-5.4 24.4 779.4 1819.1 68 3546.2 10.7-8.5 1.2 B 111.67-33.78-330.0-7.3- -9.9-20.2-490.6-1451.5-58- 2779.8-9.4-4.1-.8-116.43 40.43 1340.0 11.2-2.3 24.5 775.4 1832.2 69 3650.6 11.7 10.1 1.3 C 111.93-33.98-235.0-6.3- -11.6-21.4-488.8-1584.9-58- 2781.8-8.8-3.8-.9-117.07 40.97 920.0 10.9-2.6 24.8 746.5 1850.9 69 3647.9 11.5 8.0 1.3 D 111.67-33.77-345.0-4.4- -13.3-19.2-521.2-1369.8 59-2277.0-7.4-4.7-.7-117.13 40.95 1530.0 10.8-2.6 24.1 831.5 1797.8 71 3537.5 11.4 11.4 1.2 E 105.92-34.83-900.0-2.5- -14.7-17.0-524.4-1392.4-59- 1624.5-5.9-6.5-.7-116.77 41.03 1600.0 8.7-5.3 21.2 698.1 1661.5 65 2959.8 9.9 11.1 1.2 F 115.62-39.87-930.0-3.2- -13.9-17.7-547.7-1434.6-59- 1815.6-6.4-7.3.7 116.52 40.95 1480.0 5.3-12.6 20.7 646.5 1631.5 61 2536.5 8.1 10.4 1.0 G 115.70 39.82 680.0-6.7- -10.9-21.8-542.5-1671.8-60- 2888.4-9.0-6.0-1.0-116.55 40.38 800.0 7.6-9.7 22.6 558.8 1716.2 60 3098.1 9.6 6.7 1.1 H 116.65-40.45-810.0-6.0- -11.5-21.2-560.4-1627.5-60- 2709.7-8.6-7.0-1.0-117.13 40.53 820.0 6.2-11.5 21.4 586.6 1654.3 60 2755.5 8.7 7.4 1.0 I 112.02-35.47-920.0-3.0- -14.3-17.7 568.4-1339.4-59- 1829.4-6.3-7.5-.6-117.38 40.58 1740.0 5.7-8.6 20.8 779.1 1623.3 68 2603.2 8.3 11.9 1.0 J 111.33-33.72-1200.0-5.4- -7.4-16.9-724.7-1237.1-68- 1948.2-7.4-8.8-.5-111.95 34.60 1890.0 9.0-4.7 21.3 880.2 1531.7 71 3048.1 10.1 13.5.9 K 111.47-35.45-500.0-1.8- -15.3-15.1-511.8-1209.7-59- 1213.7-5.1-5.1-.4-117.42 41.03 2160.0 8.0-9.8 23.4 829.8 1771.6 69 3279.3 10.0 13.9 1.2 L 111.50-35.42-900.0-1.2- -16.2-15.1-550.6-1190.9-59- 1214.9-5.0-7.2-.4-116.58 40.95 2160.0 5.5-10.5 20.9 835.9 1636.9 69 2587.0 8.2 14.2 1.0 M 115.53-39.87-1280.0-3.1-13.8-17.6-588.6-1427.4-60- 1783.5-6.3-9.0-.7-115.77 40.52 1500.0 4.2-12.9 19.0 649.3 1522.7 61 2128.0 7.1 10.6.8 N 111.67-33.72-1420.0-2.2- -14.5-14.7-670.0-1165.8-62- 1277.9-5.5-10.8-.4-115.62 39.87 2230.0 8.2-4.8 19.9 884.8 1408.6 72 2742.4 9.4 14.6.7 O 115.60-39.87 1820.0-.9- -15.7-15.1-692.5-1284.8-62- 1142.9-4.6-12.4-.5-1900.0 1.3-15.3 15.6 703.6 1313.5 62 1271.3 5.0 12.8.6
1 8 8 0 500 830mm K 5 1 10 0 0 4 1 2 550 836mm K 700mm K DCA X 1 X 2 X 3 4 9 1200 1400m 1800m 30cm 50 60 30 50crn 1m ph 6 6 7 5 1760m 50cm ph 6 9 6 6 70 7 41 1g kg -1 1 9 1 1g kg -1 4.8 % CaCO 3 ph <0.01 <0.001 (H cm mm mm 2 O) g.kg -1 6 N60E 0-3 24.05 6.42 6.9 0 70.7 3 5-10 0.80 23.75 10.4 6.7 0 41.1 1.90 1 1740m 20-26 1.26 13.34 8.29 6.6 0 ---- 1.14 + 25-40 1.38 19.06 8.92 7.3 74.4 9.4 1490m 65-75 21.08 8.92 7.4 46.8 6.8 1410m 138.9g kg -1 60cm 23.8g kg -1 A 3 34cm 4.8 1.7g kg -1 1680m ph
C N 1780m C N ph C/N PH cm g.kg -1 g.kg -1 (H 2O) (HCL) (cmol.kg -1 ) (%) A6-26 A 126-42 1780m B42-76 CD76-90 A 13-10 1410m A 110-24 AB24-34 B34-60 A 14-6 1680m A 116-48 B48-74 BC74-90 136.2 82.1 33.2 32.5 138.9 77.6 41.6 23.8 186.1 67.1 30.3 --- 4.3 1.9 1.3 2.4 4.8 4.8 1.7 1.1 4.7 2.6 2.2 ---- 10.82 25.05 14.84 8.00 17.00 9.38 14.18 13.80 29.58 14.97 8.80 ---- 0.20 0.25 0.15 0.05 0.40 0.05 --- --- ---- ---- 0.05 ---- 17.5 15.0 12.5 10.0 17.5 7.5 5.0 5.0 17.5 2.0 1.0 ---- 6.8 6.5 6.6 6,2 7.1 6.5 6.7 6.5 7.0 6.3 6.7 6.7 6.8 6.5 6.2 5.2 6.8 6.0 5.9 5.7 6.6 5.2 5.3 5.3 13.49 10.28 8.40 7.66 7.06 9.26 8.77 8.15 9.17 8.42 8,52 80.92 66.84 53.85 82.37 61.44 68.39 69.62 72.19. 60.65 67.90 72.95 4.10 2000m ph 1986 1992 PH cm (H 2 O) g.kg -1 cm N50E 15 0 0-4 4-7 0-7 2450m 15-30 0.51 6.8 166.5 7-30 40-48 30-80 50-60 0.86 6.8 119.6 48-80 0-3 0-3 10 3-12 0.66 6.9 123.9 3.42 3-17 2280m 20-28 6.7 116.0 2.76 17-35 43-51 81.1 2.68 35-60 62-78 6.7 52.9 60-87 4.10 2000m
4 7 4.7 4.7 2000m
1000m 1500m 1000 1800m 1400 1900m 1900 2300m 1800m 1700 1800 m Abier sutchuenensis 2400 2800m 2400 2800m 2000m