............................................ I
....................................... 2010...... II
1989 11 89 23 :
1989 11 1992 5 28
40
15 7 9 5 5 3 6 21 25 1 2 3
1 1 2 1
65 7 9
1 2 1 2
30
10
5 3 1
1997
?
?
54mm 85mm ga98-105
1998 10 1
1. 2. 3.
40 20 10 35%
30% 25% 15% 5% 18 18 18 20% 1
1
4 1 1 2
s s s1 s2 s3 s4 s5 14 22 2 45 60% 5% 13 15 35 45%
2010 5 15 2010 2010
2000 10% 15% 50% 70%
110 20 30 6 8-10% 20
2010
110 450 bp 350 (ccic) dna
1991 19
dna
1985 18
1991 9 py py1 2 10 pw ph psh
1. 2. 3. 4. 5. 6. 7. 8. 9. 10. 1. 2. 3. 4. 5. 1. 2. 3. 4.
5. 6. 1.
2. 3. 4. 5.
1.
(ddn) 2. 420 30 3. 150 450 800 110 350
80% 4. 5.
110 119 6. 7. gps
8. 9. 10.
350 11. (gps)
(gis) 1. 2.
3. 4. 5.
6. 2000 90% 60%
7. 8. 9.