9 73852005 BullGunma MusNatuHist9 73852005 Carcharodon megalodon : 370-2345 1674-1 Carcharodon megalodon 14 12 4 Cmegalodon Carcharodon megalodon The remains of Carcharodon megalodon from the Miocene Tomioka Group Southwest Gunma Central Japan TAKAKUWA Yuji Gunma Museum of Natural History: 1674-1 Kamikuroiwa Tomioka Gunma 370-2345 JAPAN Fourteen teeth of the Megatooth shark Carcharodon megalodon are recognized from the Miocene Tomioka Group and twelve of them are reported in this paper These fossils show that the range of Cmegalodon distributed in Southwest Gunma expanded between early Middle Miocene and early Late Miocene Carcharodon megalodonsharkmiocenetomioka GroupGunma Prefecture Carcharodon megalodon Agassiz 1843 1316 Cappetta1987; Gottfried et al1996 2004 1779 1990; 2004 C.megalodon 1983 Cmegalodon 2004 19831982 NSM-PV-5432 1985 C.megalodon 1999; 2000; Takakuwa and Ando2004 2003; 2004 C megalodon CarcharodonApplegate and Espinosa-Arrubarrena 1996; Gottfried et al1996; Purdy1996 CarcharoclesCappetta1987; Kent 1994 200515200531
Map of Kanto area and Southwest Gunma showing the fossil localities Modified from Kurihara 2000 1984 1516 GMNH-PV: NSM-PV: Figure 1 199019907 Figure 22004 1990 2 1990 Cmegalodon 4 2004 1990 Cmegalodon 14 1983 212 Table 1 1983 1999 Class Chondrichthyes Subclass Elasmobranchii Bonaparte1838 Superorder Galeomorphii Compagno1973 Order Lamniformes Berg1958 Famly Lamnidae Mller et Henle1838 Genus Carcharodon Mller et Henle1838 Carcharodon megalodon Agassiz 1843 Plate Table 12
Carcharodon megalodon Stratigraphy of the Miocene Tomioka Group and fossil horizons Based upon Oishi and Takahashi1990 1989Figure 3, Table 2. 1 2 C.megalodon Figure 1Loc. 1 1-1APlate -1a,b GMNH-PV-599 Measurement points of tooth of Carcharodon megalodon (right upper tooth, modified from Uyeno et al. (1989))
List of the specimens of MegatoothsharkCarcharodon megalodon from the Miocene Tomioka Group 1214/10mm 17 1-2APlate-2 GMNH-PV-1543 10mm1112 14 2 7 2-1Plate -1a,b GMNH-PV-1588 Figure 1Loc. 2 1989 6 1982 2-2 25C.megalodon Figure 1Loc. 3 2-3BPlate-3 2GMNH-PV- 1587 2-4GMNH-PV-1544Plate-4a,b
Carcharodon megalodon 2-5Plate-2a,b GMNH-PV-1589 1971 121410mm 2-6Plate-3a,b GMNH-PV-1590 2-7Plate-4a,b,and c GMNH-PV-1591 12 14/10mm19895 3.6mm 3 3 3-1BPlate -1-1 Figure 1Loc. 4 111410mm GMNH-PV-009 3-2 NSM-PV-5432: Plate -2-2 19602231963 GMNH-PV-008 19821985 Figure 1 Loc5 1999 S 131710mm 19896 3-31983Plate -3 1983 197710 2710 1990 1983 GMNH-PV-15511 1983 plate 4 2 4-1Plate -1a,b Figure 1Loc6 157 Cmegalodon 3 10mm1820 10mm1216 19896 PV-5432
Measurementsmmof Carcharodon megalodon from the Miocene Tomioka Group 4-2Plate-2a.b Figure 1Loc. 7 GMNH-PV-010,, 10mm10121989 5,. 2004 C.megalodon 1983 3Figure 4 2004 4 C.megalodon 1983 500 1990 C.megalodon 7 5 1 Carcharodon megalodon 1412 2 C.megalodon 4
Carcharodon megalodon Range of Carcharodon megalodon in the Miocene Tomioka Group De Paul University Chicago USADr Kenshu Shimada ApplegateSPand Espinosa-Arrubarrena (1996): The fossil history of Carcharodon and its possible ancestorcretolamna In: Great White Sharks: The Biology of Carcharodon carcharias (KlimleyAPand AinleyDGeds19-36Academic PressCalifornia Cappetta H1987: ChondrichthyesMesozoic and Cenozoic ElasmobranchiiHandbook of Paleoichthyology (HPSchultze ed 3B 193ppGustav Fischer VerlagStuttgart - New York (1982): 214p (1983): Carcharodon megalodon 89: 597-598 GottfriedMDCompagnoLJVand BowmanSC(1996): Size and skeletal anatomy of the giant Megatooth shark Carcharodon megalodon In: Great White Sharks: The Biology of Carcharodon carchariasklimleyapand AinleyDGeds55-66 cademic PressCalifornia (1999): 101 KentBW(1994): Fossil Sharks of the Chesapeake Bay Region146p Egan Rees & BoyerMaryland (1779): (1971): 245p (1990): - - 92: 1-17 PurdyRW(1996): Paleoecology of fossil white sharksin: Great White Sharks: The Biology of Carcharodon carcharias KlimleyA Pand AinleyDGeds67-78Academic PressCalifornia 1990: Kanto Syntaxis 35: 51-55 2004: 1103: 175-194 TakakuwaYand AndoH2004: New selachian assemblage including deep sea sharks from the Miocene of Gunmacentral Japan Journal of Vertebrate Paleontology243supplement120A 1999: 3: 7-16 2000: 14910-11 2003:14: 62-64 2004: In : 22 8 1982: 2In : 388-389 1985: In : 74-75 1984: 2: 47-65 1989:, 7: 73-85 1999:, 321: 14-20 2004: Carcharocles megalodon : :, 75: 7-15
Plate Remains of Carcharodon megalodon from the Haratajino and Niwaya Formations 1 Left lower anterior tooth(collected by HNakajimacast as GMNH-PV-599) a lingual view; blabial view 2 Right upper lateral (8th?) tooth (collected by YSakaguchicast as GMNH-PV-1543) labial view 3 Left? lower anterior? tooth (collected by YSakaguchicast as GMNH-PV-1587) lingual view 4 Left upper posterior tooth (collected by TKurosawaGMNH-PV-1544) a lingual view; blabial view Plate Remains of Carcharodon megalodon from the Niwaya Formation 1 Right upper lateral (6th?) tooth (collected by TMoridairacast as GMNH-PV-1588) a lingual view; blabial view 2 Right? lower anterior tooth (collected by HShinozakicast as GMNH-PV-1589) a lingual view; blabial view 3 Left lower posterior? tooth (collected by KFujiicast as GMNH-PV-1590) a lingual view; blabial view 4 Left (?upper lateral (5th?) tooth (collected by YSutocast as GMNH-PV-1591) a lingual view; blabial view; cunusual keel in labial surface of apex Plate Remains of Carcharodon megalodon from the Haraichi Formation lingual view 1 Left upper anterior tooth (collected by HNakajimacast as GMNH-PV-009) 2 Left upper lateral (6th?) tooth (NSM-PV-5432cast as GMNH-PV-008) 3 Right upper tooth (cast as GMNH-PV-1551 Plate Remains of Carcharodon megalodon from the Haraichi Formation labial view 1 Left upper anterior tooth (collected by HNakajimacast as GMNH-PV-009) 2 Left upper lateral (6th?) tooth (NSM-PV-5432cast as GMNH-PV-008) Plate Remains of Carcharodon megalodon from the Itahana Formation 1Right upper lateral (6th?) tooth (Annaka City Educational BoardNo157) a lingual view; blabial view 2 Right upper lateral (5th?) tooth (collected by NFusecast as GMNH-PV-010) a lingual view; blabial view
Carcharodon megalodon Plate
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Carcharodon megalodon Plate
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Carcharodon megalodon Plate