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1 öˆ ž 3 Ò 台灣新歸化旋花科植物 - 七爪菜欒藤!!!!» * v Í r Ÿ ž 2 v Í uàæ ž 3 Ø º ºuÀ!! ì! i ˆ Merremia dissecta (Jacq.) H. Hallier. õ çý n }h Œ g p àý À à À 5-7 i ÌÁ ÅËu ç p Á k { Ç d píqq} } sg Å Á d í uÿ ªz{ˆu À! i ˆ d Åæ d Å p * ³í thhsieh@mail.nutn.edu.tw 1

ž Ì! ï é è á Í Ô «58 2000 Staples & Yang, 1998 «14 44 œ é Ð é «Í žá š ª Ž é Î ï Ipomoea L. é«13 Ž Ý Jacquemontia Choisy «1 Œ Merremia Dennst. ex Endl.

2 ž Ì! ï é è á Í Ô « Staples & Yang, 1998 «14 44 œ é Ð é «Í žá š ª Ž é Î ï Ipomoea L. é«13 Ž Ý Jacquemontia Choisy «1 Œ Merremia Dennst. ex Endl. «1 Wu, Hsieh, & Rejmanek, 2004 Œ Merremia dissecta (Jacq.) H. Hallier.) á 1979 Fang & Staples, Ÿ Y. Shimada Ð œ é Y. Shimada 3144, TAI ÐþÌ ² ÝÐ œ ô Î ž Ðþ Ô ˆ á é ÔŠ ˆ ï ž œ é žá š ª Ž é Í ŽáŠ ö ý š Õ Ì Ô Fang & Staples, 1995 é é Chang, 1978 Ô é Staples & Yang, 1998 Š¹ œ ð Î Ž éë ÈŠ «² Žë Wu, Hsieh, & Rejmanek, 2004 œ ïõÿ ï Ð ï ã 1992 ³Ÿõ É 2004 ð Î 2007 š Š œ žá ö Ð é 5-7 Ð Š öëã ïõÿ Œ Merremia Ý ï Ipomoea ë â Š á á 2007 Ð 2010 Ð ï ã 1 œ Žá ö 2

Merremia dissecta Jacquin H. Hallier, Bot. Jahrb. Syst. 16:552. 1893. Convolvulus dissectus Jacquin, Obs. Bot. 2:4. 1767.

3 Ž ï éf Œ 2/! Œ! )B*žÉ! )C* éïõ! )D* ¾lãÐ ˆë ï! h½! ï šë ï œ ž ˆ Ò ï þ á Š Îž Erdtman ë æ Erdtman, 1952 Ž þëï Îž ãè þ á ñ á ª ˆ 3000N, Hitachi Ô Œ ¹! i ˆ ³À í! Merremia dissecta Jacquin H. Hallier, Bot. Jahrb. Syst. 16: Convolvulus dissectus Jacquin, Obs. Bot. 2: Ý œ ñ ã ¾ ß ë «Žë ž é 5~7 ß ¾ Ð Š öëã Ý ß Š ñ 3~6 1~2.5 ñ 3~5 ß ë ï ž ž Ý ž ï 2.3~10 ï 2~3 Ð é 5 ä ž éß ¾ ð ˆ ñ Ð ë ã ˆ ïõ é ë 3-5 Ÿ 5 ä žáïõ ï éà«ˆý 2 ü lã ¾ 2 ü 4 5 ä ã ˆ 4 ä ï 5-10 ã 5-12 é žá Ž ò ý ãöí ˆµÍ 2:8: Ÿõ Žá ö ë! 3

粉為散孔型表面具有刺狀突起 Staples & Yang, 1998 本屬則為多種類型但表面不

5-6 溝 9-12 溝散溝及散孔型等多種 Ferguson, Verdcourt, &

microreticulate-puctate 不具刺狀突起外壁外層 sexine 為柱

4 環境與生態學報觀察標本台南市安平海拔約 5 公尺 22 Apr. 2007, 8 May 2010 NUTN, 國立臺南大學生物科技系標本室 T. H. Hsieh s. n. s. n. T. H. Hsieh 二花粉的形態本屬與牽牛花屬在外部形態上頗為相似常不易區分但是在花粉上牽牛花屬的花粉為散孔型表面具有刺狀突起 Staples & Yang, 1998 本屬則為多種類型但表面不具刺狀突起根據花粉資料可以支持將本種歸類於菜欒藤屬菜欒藤屬的花粉有多種類型分別為 3 溝 5-6 溝 9-12 溝散溝及散孔型等多種 Ferguson, Verdcourt, & Poole, 1977 本種花粉為 3 溝粒屬於第一種類型表面具有細網紋至細穿孔紋 microreticulate-puctate 不具刺狀突起外壁外層 sexine 為柱狀結構具蓋頂層圖 2 10μm 10μm 1μm 10μm 1μm 圖七爪菜欒藤之花粉掃描式電子顯微鏡照像圖 2. A.赤道面溝間區 B.赤道面溝區 C.極面觀 D.外壁紋飾為細網紋至細穿孔紋 E.花粉外壁為柱狀結構具蓋頂層 4

5 Ž ï éf Œ u ³! Œ žñá žé Ý œ é ö ž ñ Ì ï ã žñ Î Ã Ð ²žÐ É é l Tribulus taiwanense T. C. Huang et T. H. Hsieh l Cenchrus echinatus L. Œ Chloris barbata Sw. ˆ Panicum maximum Jacq. Mimosa pudica L. ê Cynodon dactylon (L.) Pers. ˆ ï Bidens pilosa L. var. radiata Sch. òˆ Cyperus rotundus L. Ê Alysicarpus vaginalis (L.) DC. Chenopodium acuminatum Willd. ssp. virgatum (Thunb.) Kitamura Chamaesyce hirta (L.) Millsp. Ÿ Sesbania cannabiana (Retz.) Poir Rhynchelytrum repens Willd. C. E. Hubb. ê¼ Heliotropium indicum L. Ý œ é! œ ð Þ 2 ± í Ì Î ë É œ ÈÞ Ö Î È G1 5

6 ž g! 1979 Š é¾ 64(1): éë Í 1992 é éë 2004 œ ï éë Š ˆ 1976 šî é éë Chang, C.E. (1978). Convolvulaceae. In H.L. Li et al. (eds.) Flora of Taiwan 4, Epoch Publ. Co., Ltd, Taipei. Erdtman, G. (1952). Pollen Morphology and Plant Taxonomy. The Chronica Botanica Co. Waltham, Mass., USA. Ferguson, I.K., Verdcourt, T. & Poole, M.M. (1977). Pollen morphology in the genera Merremia and Operculina (Convolvulaceae) and its taxonomic significance. Kew Bull. 31(4), Fang, R.C. & Staples, G. W. (1995). Convolvulaceae. In Z.Y. Wu and P.H. Raven (eds.), Flora of China 16, Staples, G.W. &. Yang, S. Z. (1998). Convolvulaceae. In T.C. Huang et al. (eds.) Flora of Taiwan 2 nd ed. 4, Dept. Bot. NTU. Taipei. Wu, S.H., Hsieh, C.F. & Rejmanek, M. (2004). Catalogue of the naturalized flora of Taiwan. Taiwania 49, Á Ò

7 Ž ï éf Œ Merremia dissecta (Jacq.) H. Hallier. A newly naturalized species in Taiwan Yu-Chao Tsai 1 Pau-Jee Lee 2 Yi-Han Chang 3 and Tsung-Hsin Hsieh 2* 1 Department of Early Childhood Education, National University of Tainan, Tainan, Taiwan Department of Biological Sciences and Technology, National University of Tainan, Tainan, Taiwan 3 Division of Forest Biology, Taiwan Forestry Research Institute, Taipei, Taiwan. Abstract Alamo vine (Merremia dissecta (Jacq.) H. Hallier.), newly naturalized in Taiwan, is native to tropical America. It was possibly introduced and cultivated as garden plant during the period of Japanese Occupation of Taiwan. This species can be distinguished from other relative species on the island by its 5-7-divided palmate leaves, coarsely dentate to irregularly pinnately parted on the margin, and white corolla with purple-red on the throat. In our recent plant inventory, a wild population was discovered at Anping area, Tainan City, Southern Taiwan. A detailed description with color photographs, pollen s SEM microphotographs and habitat information are provided in this paper. Key words: Merremia dissecta, naturalized plant, Convolvulaceae, taxonomy, pollen, Taiwan * Corresponding author: thhsieh@mail.nutn.edu.tw 7

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